Sphenosuchia
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Sphenosuchia
Sphenosuchia is a suborder of basal crocodylomorphs that first appeared in the Triassic and occurred into the Middle Jurassic. Most were small, gracile animals with an erect limb posture. They are now thought to be ancestral to crocodyliforms, a group which includes all living crocodilians. Stratigraphic range The earliest known members of the group (i.e. '' Hesperosuchus'') are early Norian in age, found in the Blue Mesa Member of the Chinle Formation. Only one sphenosuchian is currently known from the Middle Jurassic, ''Junggarsuchus'', from the Junggar Basin (Shishugou Formation) of China during either the Bathonian or the Callovian (~165 Ma) age,Clark, J.M., ''et al.'' (2004A Middle Jurassic 'sphenosuchian' from China and the origin of the crocodylian skull''Nature'' 430:1021-1024. and the Hallopodidae are known from the Late Jurassic of North America. Phylogeny The monophyly of the group is debated, although several synapomorphies characterize the clade, including extreme ...
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Saltoposuchus
''Saltoposuchus'' is an extinct genus of small (1-1.5 m and 10-15 kg), long-tailed crocodylomorph reptile (Sphenosuchia), from the Norian (Late Triassic) of Europe. The name translated means "leaping foot crocodile". It has been proposed that ''Terrestrisuchus gracilis'' and ''Saltoposuchus connectens'' represent different ontogenetic stages of the same genus. ''Saltoposuchus'' was commonly (and incorrectly) referred to in popular literature as the ancestor (or close ancestors) to dinosaurs, however, recent scientific research show that this is not the case. Description and Paleobiology Fossil evidence of Sphenosuchia and early crocodylomorphs lead paleontologists to conclude that ''Saltoposuchus'' is a terrestrial animal. As a monophyletic group of crocodylomorphs, Saltoposuchidae have many key morphological traits shared with most crocodylomorphs. Skull Much like other crocodylomorphs, ''Saltoposuchus'' skulls had a (reduced) antorbital fenestra, an overhanging squa ...
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Sphenosuchia
Sphenosuchia is a suborder of basal crocodylomorphs that first appeared in the Triassic and occurred into the Middle Jurassic. Most were small, gracile animals with an erect limb posture. They are now thought to be ancestral to crocodyliforms, a group which includes all living crocodilians. Stratigraphic range The earliest known members of the group (i.e. '' Hesperosuchus'') are early Norian in age, found in the Blue Mesa Member of the Chinle Formation. Only one sphenosuchian is currently known from the Middle Jurassic, ''Junggarsuchus'', from the Junggar Basin (Shishugou Formation) of China during either the Bathonian or the Callovian (~165 Ma) age,Clark, J.M., ''et al.'' (2004A Middle Jurassic 'sphenosuchian' from China and the origin of the crocodylian skull''Nature'' 430:1021-1024. and the Hallopodidae are known from the Late Jurassic of North America. Phylogeny The monophyly of the group is debated, although several synapomorphies characterize the clade, including extreme ...
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Litargosuchus
''Litargosuchus'' is a sphenosuchian Crocodylomorpha, crocodylomorph, a Basal (phylogenetics), basal member of the Crocodylomorpha, crocodylomorph clade from the Early Jurassic of South Africa. Its genus name ''Litargosuchus'' is derived from Greek meaning "fast running crocodile" and its species name ''leptorhynchus'' refers to its Gracility, gracile snout. ''Litargosuchus'', along with all of South Africa's Crocodylomorpha, crocodylomorph Taxon, taxa, are confined to the upper Elliot Formation (UEF) in South Africa. History of discovery In 1988 James Kitching found the holotype fossil of ''Litargosuchus'' in a field expedition in Eagles Crag, a farm near Barkly East in the Eastern Cape. The material recovered consisted of a near-complete skull with its mandible and several bones belonging to the postcranial skeleton. At the time, James Kitching, Kitching was the director of the Evolutionary Studies Institute (previously the Bernard Price Institute) of the University of the Witw ...
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Terrestrisuchus
''Terrestrisuchus'' is an extinct genus of very small early crocodylomorph that was about long. Fossils have been found in Wales and Southern England and date from near the very end of the Late Triassic during the Rhaetian, and it is known by type and only known species ''T. gracilis''. ''Terrestrisuchus'' was a long-legged, active predator that lived entirely on land, unlike modern crocodilians. It inhabited a chain of tropical, low-lying islands that made up southern Britain, along with similarly small-sized dinosaurs and abundant rhynchocephalians. Numerous fossils of ''Terrestrisuchus'' are known from fissures in limestone karst which made up the islands it lived on, which formed caverns and sinkholes that preserved the remains of ''Terrestrisuchus'' and other island-living reptiles. Description ''Terrestrisuchus'' was a small, slender crocodylomorph with very long legs, quite unlike modern crocodilians. It was initially estimated to have been between long, although this e ...
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Crocodylomorph
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, crocodylomorphs were far more diverse than they are now. Triassic forms were small, lightly built, active terrestrial animals. The earliest and most primitive crocodylomorphs are represented by " sphenosuchians", a paraphyletic assemblage containing small-bodied forms with elongated limbs that walked upright, which represents the ancestral morphology of Crocodylomorpha. These forms persisted until the end of the Jurassic. During the Jurassic, Crocodylomorphs morphologically diversified into numerous niches, including into the aquatic and marine realms. Evolutionary history When their extinct species and stem group are examined, the crocodylian lineage (clade Pseudosuchia, formerly Crurotarsi) proves to have been a very diverse and adaptive ...
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Kayentasuchus
''Kayentasuchus'' (meaning "Kayenta Formation crocodile") is a genus of sphenosuchian, a type of basal crocodylomorph, the clade that comprises the crocodilians and their closest kin. It is known from a single skeleton found in rocks of the Sinemurian-Pliensbachian-age Lower Jurassic Kayenta Formation, northeastern Arizona. History and description ''Kayentasuchus'' is based on UCMP 131830, a fragmentary skeleton. UCMP 131830 was known for many years before its description as the "Kayenta Form". It was found in 1983 by James M. Clark. The location was at Willow Springs, northeast of Flagstaff. The specimen was found in a channel sandstone about halfway up the formation, in the "silty facies". Specimens of the turtle '' Kayentachelys'', the dinosaur ''Scutellosaurus'', and tritylodonts were found in the immediate vicinity a few meters below it. ''Kayentasuchus'' was named in 2002 by Clark and Hans-Dieter Sues. The type species is ''K. walkeri'' in memory of ...
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Hesperosuchus
''Hesperosuchus'' is an extinct genus of crocodylomorph reptile that contains a single species, ''Hesperosuchus agilis''. Remains of this pseudosuchian have been found in Late Triassic (Carnian) strata from Arizona and New Mexico.Colbert, E. H. 1952. A pseudosuchian reptile from Arizona. Bulletin of the American Museum of Natural History 99:561–592. Because of similarities in skull and neck anatomy and the presence of hollow bones ''Hesperosuchus'' was formerly thought to be an ancestor of later carnosaurian dinosaurs, but based on more recent findings and research it is now known to be more closely related to crocodilians rather than dinosaurs.Brinkman, D. 1981. The origin of the crocodiloid tarsi and the interrelationships of thecodontian archosaurs. Breviora 464:1–23.Benton, M. J. and J. M. Clark. 1988. Archosaur phylogeny and the relationships of the Crocodylia. pp. 295–338. In M. J. Benton (ed.). The Phylogeny and Classification of the Tetrapods Vol. 1.Amphibians, Reptil ...
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Pseudhesperosuchus
''Pseudhesperosuchus'' (meaning "false '' Hesperosuchus''") is a genus of sphenosuchian, a type of basal crocodylomorph, the clade that comprises the crocodilians and their closest kin. It is known from a partial skeleton and skull found in rocks of the Late Triassic ( Norian-age) Los Colorados Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina. History and description ''Pseudhesperosuchus'' is based on PVL 3830. This specimen consists of a skull and lower jaws, most of the vertebral column, the shoulder girdle, and parts of the arms and legs. The genus was named by José Bonaparte in 1969. The type species is ''P. jachaleri''. The skull, though nearly complete, is poorly preserved, and some of its bones and sutures have been misidentified over the years. A 2002 phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, b ...
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Sphenosuchus
''Sphenosuchus'' ("wedge crocodile") is an extinct genus of crocodylomorph from the Early Jurassic Elliot Formation of South Africa, discovered and described early in the 20th century. The skull is preserved very well but other than elements of the forelimb and isolated parts of the hind limb, the ''Sphenosuchus'' material is incomplete. It was probably quadrupedal, but may have been a facultative biped. ''Sphenosuchus'' was first thoroughly described in 1972 by the British palaeontologist Alick Walker, in a paper in the journal ''Nature''. Walker suggested, based on detailed (but still preliminary at that time) studies of the skull of ''Sphenosuchus'' and modern birds, that crocodylomorphs and birds might share an immediate common ancestor. Walker recanted his hypothesis in 1985, but restated and elaborated on it (in essence 'de-recanting') in a monograph published in 1990, which provided the most comprehensive description and discussion of ''Sphenosuchus'' yet published. Bro ...
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Dibothrosuchus
''Dibothrosuchus'' is a genus of sphenosuchian, a type of basal crocodylomorph, the clade that comprises the crocodilians and their closest kin. It is known from several partial skeletons and skulls. These fossils were found in Lower Jurassic (Lower Pliensbachian, circa 186 million years old) rocks of Yunnan, China. ''Dibothrosuchus'' was a small terrestrial crocodylomorph that probably had a keen sense of hearing, and thus was probably a vocal animal like modern crocodilians. History and description ''Dibothrosuchus'' was named in 1965 by D.J. Simmons based on a partial skull and skeleton collected by Father Oehler of Fu Jen Catholic University from mudstones near Huangchiatien. The type species is ''D. elaphros''. Additional remains of this genus were recovered in 1985 by a joint Chinese-US expedition. Fossils of ''Dibothrosuchus'' come from the Zhangjiawa Formation, being originally assigned to the Reds Beds of the Lufeng Formation, thus being geologically young ...
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Synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Monophyly
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have taken ...
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