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Stylonurine
Stylonurina is one of two suborders of eurypterids, a group of extinct arthropods commonly known as "sea scorpions". Members of the suborder are collectively and informally known as "stylonurine eurypterids" or "stylonurines". They are known from deposits primarily in Europe and North America, but also in Siberia. Compared to the other suborder, Eurypterina, the stylonurines were comparatively rare and retained their posterior prosomal appendages for walking. Despite their rarity, the stylonurines have the longest temporal range of the two suborders. The suborder contains some of the oldest known eurypterids, such as ''Brachyopterus'', from the Middle Ordovician as well as the youngest known eurypterids, from the Late Permian. They remained rare throughout the Ordovician and Silurian, though the radiation of the mycteropoids (a group of large sweep-feeding forms) in the Late Devonian and Carboniferous is the last major radiation of the eurypterids before their extinction in the Pe ...
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Eurypterid
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 Myr, million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic Chelicerata, chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Tri ...
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Pagea
''Pagea'' is a genus of prehistoric eurypterid classified as part of the family Stylonuridae. It contains three species, all from the Devonian (Lochkovian to Pragian); ''P. plotnicki'' from Nunavut, Canada and ''P. sturrocki'' and ''P. symondsii'' from the Old Red Sandstone of the United Kingdom.Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 16.0 http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf (PDF). The genus is named in honor of David Page, an early worker on the fauna of the Old Red Sandstone and describer of the first Stylonurine eurypterid. Description ''Pagea'' was a large stylonurid eurypterid. The third and fourth prosomal appendages bore double rows of flat spines. The prosoma was subrectangularly shaped, with the eyes located on the anterior half. The metastoma was narrow in relation to the width of the prosoma, ...
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Rhenopteroidea
The Rhenopteridae are a family of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". The family is the only family currently contained in the superfamily Rhenopteroidea, one of four superfamilies classified as part of the suborder Stylonurina. The family contains one of the earliest known eurypterids, '' Brachyopterus'', known from the Middle Ordovician (also the sister taxon to all other rhenopterids), and is the most primitive clade of stylonurine eurypterids. The last known members of the family went extinct during the Early Devonian. Description Rhenopterids were small, characterized by scattered tubercules and knobs on the outer surface of the exoskeleton. Their first two (or possibly three) pairs of walking legs had spines; the last two pairs were long and powerful, without spines. The prosoma (head) was subtrapezoidal, with arcuate compound eyes on parallel axes. The male genital appendages were short with two distal spines. The ...
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Mycteropoid
Mycteropoidea is an extinct superfamily of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". It is one of four superfamilies classified as part of the suborder Stylonurina. Mycteropoids have been recovered from Europe, Russia, South America and South Africa. Mycteropoid specimens are often fragmentary, making it difficult to establish relationships between the included taxa. Only two mycteropoid taxa are known from reasonable complete remains, ''Hibbertopterus scouleri'' and ''H. wittebergensis''. Mycteropoids were large bizarre Eurypterids found from the Early Silurian to the end of the Permian period. They were sweep feeders, inhabiting freshwater swamps and rivers, feeding by raking through the soft sediment with blades on their anterior appendages to capture small invertebrates. Their morphology was so unusual that they have been thought to be an order separate to Eurypterida. Recent work however confirms them to be derived members of t ...
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Stylonurella
''Stylonurella'' is a genus of prehistoric eurypterid. It is classified within the Parastylonuridae family and contains three species, ''S. arnoldi'' and ''S. beecheri'' from the Devonian of Pennsylvania, United States and ''S. spinipes'' from the Silurian of Kip Burn, Scotland.Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 16.0 http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf (PDF). Description ''Stylonurella'' was a small stylonuroid, possessing a subquadrate prosoma with approximately the same length as width. The midsection was slightly constricted and the eyes were parallel and anteriorly located in the anterior half of the carapace. The metastoma and first two appendages are unknown, the third and fourth prosomal legs are very short and the last two walking legs are very long. The metasoma is very narrow. A revision of th ...
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Stylonurus
''Stylonurus'' is a genus of prehistoric eurypterid of the family Stylonuridae. The genus contains three species: ''Stylonurus powriensis'' from the Devonian of Scotland, ''Stylonurus shaffneri'' from the Devonian of Pennsylvania and ''Stylonurus perspicillum'' from the Devonian of Germany.Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 16.0 http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf (PDF). The assignment of ''S. perspicillum'' and ''S. shaffneri'' to the genus is doubtful. A previously assigned species, ''S. ensiformis'', is today regarded as synonymous with ''S. powriensis''. Description Stylonurids, which lived from the Ordovician to Lower Permian periods, were small to very large forms with scales developing into tubercules and knobs. The prosoma (head) exhibited variable shape, with arcuate compound eyes located subcen ...
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Rhenopterus
''Rhenopterus'' is an extinct prehistoric eurypterid. Fossils of ''Rhenopterus'' have been recovered from deposits of Lower Devonian age in Germany.Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 16.0 http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf (PDF). Description Rhenopterids were small, characterized by scattered tubercules and knobs on the outer surface of the exoskeleton. Their first two (or possibly three) pairs of walking legs had spines; the last two pairs were long and powerful, without spines. The prosoma (head) was subtrapezoidal, with arcuate compound eyes on parallel axes. The male genital appendages were short with two distal spines. Species ''Rhenopterus'' contains three valid species, with other named species now seen as invalid or part of other genera. * ''Rhenopterus diensti'' Størmer, 1936 - Germany (Lower D ...
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Kiaeropterus
''Kiaeropterus'' is a genus of prehistoric eurypterid classified as part of the Rhenopteridae family. Two species, both from the Silurian period, are known; ''K. cyclophthalmus'' from Scotland and ''K. ruedemanni'' from Norway.Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 16.0 http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf (PDF). Description ''Kiaeropterus'' was a small eurypterid, with ''K. cyclophthalmus'' reaching lengths of about 6,6 cm. It had a subquadrate carapace with broad marginal rims. The eyes were large, covering about a third of the length of the carapace. The metasoma was narrow and the mesosoma not wider than the carapace, unlike in some eurypterids. The appendages V and VI were non-spiniferous and simple with individual podomeres having pronounced ridges. The genus is very poorly known. The type spec ...
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The Eurypterida Of New York Figure 63
''The'' () is a grammatical article in English, denoting persons or things that are already or about to be mentioned, under discussion, implied or otherwise presumed familiar to listeners, readers, or speakers. It is the definite article in English. ''The'' is the most frequently used word in the English language; studies and analyses of texts have found it to account for seven percent of all printed English-language words. It is derived from gendered articles in Old English which combined in Middle English and now has a single form used with nouns of any gender. The word can be used with both singular and plural nouns, and with a noun that starts with any letter. This is different from many other languages, which have different forms of the definite article for different genders or numbers. Pronunciation In most dialects, "the" is pronounced as (with the voiced dental fricative followed by a schwa) when followed by a consonant sound, and as (homophone of the archaic ...
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Hibbertopterus Scouleri
''Hibbertopterus'' is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of ''Hibbertopterus'' have been discovered in deposits ranging from the Devonian period in Belgium, Scotland and the United States to the Carboniferous period in Scotland, Ireland, the Czech Republic and South Africa. The type species, ''H. scouleri'', was first named as a species of the significantly different ''Eurypterus'' by Samuel Hibbert in 1836. The generic name ''Hibbertopterus'', coined more than a century later, combines his name and the Greek word πτερόν (''pteron'') meaning "wing". ''Hibbertopterus'' was the largest eurypterid within the stylonurine suborder, with the largest fossil specimens suggesting that ''H. scouleri'' could reach lengths around 180–200 centimetres (5.9–6.6 ft). Though this is significantly smaller than the largest eurypterid overall, ''Jaekelopterus'', which could reach lengths of around , ''Hibbertopterus'' is likely to have been the h ...
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Jaekelopterus
''Jaekelopterus'' is a genus of predatory eurypterid, a group of extinct aquatic arthropods. Fossils of ''Jaekelopterus'' have been discovered in deposits of Early Devonian age, from the Pragian and Emsian stages. There are two known species: the type species ''J. rhenaniae'' from brackish to fresh water strata in the Rhineland, and ''J. howelli'' from estuarine strata in Wyoming. The generic name combines the name of German paleontologist Otto Jaekel, who described the type species, and the Greek word πτερόν (''pteron'') meaning "wing". Based on the isolated fossil remains of a large chelicera (claw) from the Klerf Formation of Germany, ''J. rhenaniae'' has been estimated to have reached a size of around 2.3–2.6 metres (7.5–8.5 ft), making it the largest arthropod ever discovered, surpassing other large arthropods such as fellow eurypterids '' Acutiramus'' and ''Pterygotus'' and the millipede ''Arthropleura''. ''J. howelli'' was much smaller, reaching 80 ce ...
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Pterygotus
''Pterygotus'' is a genus of giant predatory eurypterid, a group of extinct aquatic arthropods. Fossils of ''Pterygotus'' have been discovered in deposits ranging in age from Middle Silurian to Late Devonian, and have been referred to several different species. Fossils have been recovered from four continents; Australia, Europe, North America and South America, which indicates that ''Pterygotus'' might have had a nearly cosmopolitan (worldwide) distribution. The type species, ''P. anglicus'', was described by Swiss naturalist Louis Agassiz in 1839, who gave it the name ''Pterygotus'', meaning "winged one". Agassiz mistakenly believed the remains were of a giant fish; he would only realize the mistake five years later in 1844. ''Pterygotus'' was among the largest eurypterids. Isolated fossil remains of a large chelicera (frontal appendage) suggests that the largest known species, ''P. grandidentatus'', reached a body length of . Several other species, notably ''P. impacatus'' at ...
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