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Shade Avoidance
Shade avoidance is a set of responses that plants display when they are subjected to the shade of another plant. It often includes elongation, altered flowering time, increased apical dominance and altered partitioning of resources. This set of responses is collectively called the shade-avoidance syndrome (SAS). Shade responses display varying strength along a continuum. Most plants are neither extreme shade avoiders or tolerators, but possess a combination of the two strategies; this helps adapt them to their environment. However, the ability to perceive and respond to shade plays a very important role in all plants: they are sessile by nature and access to photosynthetically active radiation is essential for plant nutrition and growth. In addition, the time at which a plant starts to flower is affected by the amount of light that is available. Over the past few decades, major increases in grain yield have come largely through increasing planting densities. As planting densiti ...
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Shade (shadow)
Shade is the blocking of sunlight (in particular direct sunshine) by any object, and also the shadow created by that object. Shade also consists of the colors grey, black, white, etc. It may refer to blocking of sunlight by a roof, a tree, an umbrella, a window shade or blind, curtains, or other objects. As a resource In temperate and tropical zones (most places on Earth), shade is an important issue in providing cooling and shelter from harmful heat and ultraviolet emitted by the Sun. Plants Green plants produce shade by absorbing sunlight to invest as energy in photosynthesis to produce sugar. They also actively transpire, producing an additional cooling effect. In gardening terms, there are various types of shade: *Full sun – more than five hours of direct sun per day. *Part shade – two to five hours of direct sun, or all-day ' (sunlight shining through open trees). *Full shade – less than two hours of direct sun per day... Under a dense forest canopy, light in ...
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Hypocotyl
The hypocotyl (short for "hypocotyledonous stem", meaning "below seed leaf") is the stem of a germinating seedling, found below the cotyledons (seed leaves) and above the radicle (root). Eudicots As the plant embryo grows at germination, it sends out a shoot called a radicle that becomes the primary root, and then penetrates down into the soil. After emergence of the radicle, the hypocotyl emerges and lifts the growing tip (usually including the seed coat) above the ground, bearing the embryonic leaves (called cotyledons), and the plumule that gives rise to the first true leaves. The hypocotyl is the primary organ of extension of the young plant and develops into the stem. Monocots The early development of a monocot seedling like cereals and other grasses is somewhat different. A structure called the coleoptile, essentially a part of the ''cotyledon'', protects the young stem and plumule as growth pushes them up through the soil. A mesocotyl—that part of the young plant that li ...
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Gene Redundancy
Gene redundancy is the existence of multiple genes in the genome of an organism that perform the same function. Gene redundancy can result from gene duplication. Such duplication events are responsible for many sets of paralogous genes. When an individual gene in such a set is disrupted by mutation or targeted knockout, there can be little effect on phenotype as a result of gene redundancy, whereas the effect is large for the knockout of a gene with only one copy. Gene knockout is a method utilized in some studies aiming to characterize the maintenance and fitness effects functional overlap. Classical models of maintenance propose that duplicated genes may be conserved to various extents in genomes due to their ability to compensate for deleterious loss of function mutations.Nowak MA, Boerlijst MC, Cooke J, Smith JM. 1997. nowak_smith_1997_evolution_of_genetic_redundancy_Nature97. 275:1–5. sftp://cerca@192.168.2.5/home/cerca/Desktop/data/laptop_files/info/biologia/filogeny_evoluti ...
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Brassinosteroid
Brassinosteroids (BRs or less commonly BS) are a class of polyhydroxysteroids that have been recognized as a sixth class of plant hormones and may have utility as an anticancer drug for endocrine-responsive cancers to induce apoptosis and inhibit growth. These brassinosteroids were first explored during the 70s, when Mitchell et al. reported promotion in stem elongation and cell division by the treatment of organic extracts of rapeseed ('' Brassica napus'') pollen. Brassinolide was the first isolated brassinosteroid in 1979, when pollen from '' Brassica napus'' was shown to promote stem elongation and cell divisions, and the biologically active molecule was isolated. The yield of brassinosteroids from 230 kg of ''Brassica napus'' pollen was only 10 mg. Since their discovery, over 70 BR compounds have been isolated from plants. Biosynthesis The BR is biosynthesised from campesterol. The biosynthetic pathway was elucidated by Japanese researchers and later shown to ...
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Genetic Variation
Genetic variation is the difference in DNA among individuals or the differences between populations. The multiple sources of genetic variation include mutation and genetic recombination. Mutations are the ultimate sources of genetic variation, but other mechanisms, such as genetic drift, contribute to it, as well. Among individuals within a population Genetic variation can be identified at many levels. Identifying genetic variation is possible from observations of phenotypic variation in either quantitative traits (traits that vary continuously and are coded for by many genes (e.g., leg length in dogs)) or discrete traits (traits that fall into discrete categories and are coded for by one or a few genes (e.g., white, pink, or red petal color in certain flowers)). Genetic variation can also be identified by examining variation at the level of enzymes using the process of protein electrophoresis. Polymorphic genes have more than one allele at each locus. Half of the genes ...
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Reproductive Success
Reproductive success is an individual's production of offspring per breeding event or lifetime. This is not limited by the number of offspring produced by one individual, but also the reproductive success of these offspring themselves. Reproductive success is different from fitness in that individual success is not necessarily a determinant for adaptive strength of a genotype since the effects of chance and the environment have no influence on those specific genes. Reproductive success turns into a part of fitness when the offspring are actually recruited into the breeding population. If offspring quantity is not correlated with quality this holds up, but if not then reproductive success must be adjusted by traits that predict juvenile survival in order to be measured effectively. Quality and quantity is about finding the right balance between reproduction and maintenance. The disposable soma theory of aging tells us that a longer lifespan will come at the cost of reproduction a ...
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Axillary Bud
The axillary bud (or lateral bud) is an embryonic or organogenic shoot located in the axil of a leaf. Each bud has the potential to form shoots, and may be specialized in producing either vegetative shoots (stems and branches) or reproductive shoots (flowers). Once formed, a bud may remain dormant for some time, or it may form a shoot immediately. Overview An axillary bud is an embryonic or organogenic shoot which lies dormant at the junction of the stem and petiole of a plant. It arises exogenously from outer layer of cortex of the stem. Axillary buds do not become actively growing shoots on plants with strong apical dominance (the tendency to grow just the terminal bud on the main stem). Apical dominance occurs because the shoot apical meristem produces auxin which prevents axillary buds from growing. The axillary buds begin developing when they are exposed to less auxin, for example if the plant naturally has weak apical dominance, if apical dominance is broken by removi ...
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Rosette (botany)
In botany, a rosette is a circular arrangement of leaves or of structures resembling leaves. In flowering plants, rosettes usually sit near the soil. Their structure is an example of a modified stem in which the internode gaps between the leaves do not expand, so that all the leaves remain clustered tightly together and at a similar height. Some insects induce the development of galls that are leafy rosettes. In bryophytes and algae, a rosette results from the repeated branching of the thallus as the plant grows, resulting in a circular outline. Taxonomies Many plant families have varieties with rosette morphology; they are particularly common in Asteraceae (such as dandelions), Brassicaceae (such as cabbage), and Bromeliaceae. The fern '' Blechnum fluviatile'' or New Zealand Water Fern (''kiwikiwi'') is a rosette plant. Function in flowering plants Often, rosettes form in perennial plants whose upper foliage dies back with the remaining vegetation protecting the plant. Ano ...
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Petiole (botany)
In botany, the petiole () is the stalk that attaches the leaf blade to the stem, and is able to twist the leaf to face the sun. This gives a characteristic foliage arrangement to the plant. Outgrowths appearing on each side of the petiole in some species are called stipules. Leaves with a petiole are said to be petiolate, while leaves lacking a petiole are called sessile or apetiolate. Description The petiole is a stalk that attaches a leaf to the plant stem. In petiolate leaves, the leaf stalk may be long, as in the leaves of celery and rhubarb, or short. When completely absent, the blade attaches directly to the stem and is said to be sessile. Subpetiolate leaves have an extremely short petiole, and may appear sessile. The broomrape family Orobanchaceae is an example of a family in which the leaves are always sessile. In some other plant groups, such as the speedwell genus '' Veronica'', petiolate and sessile leaves may occur in different species. In the grasses (Poaceae), ...
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Dephosphorylation
In biochemistry, dephosphorylation is the removal of a phosphate (PO43−) group from an organic compound by hydrolysis. It is a reversible post-translational modification. Dephosphorylation and its counterpart, phosphorylation, activate and deactivate enzymes by detaching or attaching phosphoric esters and anhydrides. A notable occurrence of dephosphorylation is the conversion of ATP to ADP and inorganic phosphate. Dephosphorylation employs a type of hydrolytic enzyme, or hydrolase, which cleaves ester bonds. The prominent hydrolase subclass used in dephosphorylation is phosphatase, which removes phosphate groups by hydrolysing phosphoric acid monoesters into a phosphate ion and a molecule with a free hydroxyl (-OH) group. The reversible phosphorylation-dephosphorylation reaction occurs in every physiological process, making proper function of protein phosphatases necessary for organism viability. Because protein dephosphorylation is a key process involved in cell signallin ...
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Auxin
Auxins (plural of auxin ) are a class of plant hormones (or plant-growth regulators) with some morphogen-like characteristics. Auxins play a cardinal role in coordination of many growth and behavioral processes in plant life cycles and are essential for plant body development. The Dutch biologist Frits Warmolt Went first described auxins and their role in plant growth in the 1920s. Kenneth V. Thimann became the first to isolate one of these phytohormones and to determine its chemical structure as indole-3-acetic acid (IAA). Went and Thimann co-authored a book on plant hormones, ''Phytohormones'', in 1937. Overview Auxins were the first of the major plant hormones to be discovered. They derive their name from the Greek word αυξειν (''auxein'' – "to grow/increase"). Auxin is present in all parts of a plant, although in very different concentrations. The concentration in each position is crucial developmental information, so it is subject to tight regulation through both meta ...
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Rosette (botany)
In botany, a rosette is a circular arrangement of leaves or of structures resembling leaves. In flowering plants, rosettes usually sit near the soil. Their structure is an example of a modified stem in which the internode gaps between the leaves do not expand, so that all the leaves remain clustered tightly together and at a similar height. Some insects induce the development of galls that are leafy rosettes. In bryophytes and algae, a rosette results from the repeated branching of the thallus as the plant grows, resulting in a circular outline. Taxonomies Many plant families have varieties with rosette morphology; they are particularly common in Asteraceae (such as dandelions), Brassicaceae (such as cabbage), and Bromeliaceae. The fern '' Blechnum fluviatile'' or New Zealand Water Fern (''kiwikiwi'') is a rosette plant. Function in flowering plants Often, rosettes form in perennial plants whose upper foliage dies back with the remaining vegetation protecting the plant. Ano ...
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