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Rami Communicantes
Ramus communicans (plural rami communicantes) is the Latin term used for a nerve which connects two other nerves, and can be translated as "communicating branch". Structure When used without further definition, it almost always refers to a communicating branch between a spinal nerve and the sympathetic trunk. More specifically, it usually refers to one of the following : * Gray ramus communicans * White ramus communicans The grey and white rami communicantes are responsible for conveying autonomic signals, specifically for the sympathetic nervous system. Their difference in coloration is caused by differences in myelination of the nerve fibres contained within, i.e. there are more myelinated than unmyelinated fibres in the white rami communicantes while the converse is true for the grey rami communicantes. Gray ramus communicans The grey rami communicantes exist at every level of the spinal cord and are responsible for carrying postganglionic nerve fibres from the paravertebr ...
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Spinal Nerve
A spinal nerve is a mixed nerve, which carries motor, sensory, and autonomic signals between the spinal cord and the body. In the human body there are 31 pairs of spinal nerves, one on each side of the vertebral column. These are grouped into the corresponding cervical, thoracic, lumbar, sacral and coccygeal regions of the spine. There are eight pairs of cervical nerves, twelve pairs of thoracic nerves, five pairs of lumbar nerves, five pairs of sacral nerves, and one pair of coccygeal nerves. The spinal nerves are part of the peripheral nervous system. Structure Each spinal nerve is a mixed nerve, formed from the combination of nerve fibers from its dorsal and ventral roots. The dorsal root is the afferent sensory root and carries sensory information to the brain. The ventral root is the efferent motor root and carries motor information from the brain. The spinal nerve emerges from the spinal column through an opening (intervertebral foramen) between adjacent vertebrae. ...
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Sympathetic Trunk
The sympathetic trunks (sympathetic chain, gangliated cord) are a paired bundle of nerve fibers that run from the base of the skull to the coccyx. They are a major component of the sympathetic nervous system. Structure The sympathetic trunk lies just lateral to the vertebral bodies for the entire length of the vertebral column. It interacts with the anterior rami of spinal nerves by way of rami communicantes. The sympathetic trunk permits preganglionic fibers of the sympathetic nervous system to ascend to spinal levels superior to T1 and descend to spinal levels inferior to L2/3.Greenstein B., Greenstein A. (2002): Color atlas of neuroscience – Neuroanatomy and neurophysiology. Thieme, Stuttgart – New York, . The superior end of it is continued upward through the carotid canal into the skull, and forms a plexus on the internal carotid artery; the inferior part travels in front of the coccyx, where it converges with the other trunk at a structure known as the ganglion impar. ...
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Gray Ramus Communicans
Each spinal nerve receives a branch called a gray ramus communicans (plural rami communicantes) from the adjacent paravertebral ganglion of the sympathetic trunk. The gray rami communicantes contain postganglionic nerve fibers of the sympathetic nervous system and are composed of largely unmyelinated neurons. This is in contrast to the white rami communicantes, in which heavily myelinated neurons give the rami their white appearance. Function Preganglionic sympathetic fibres from the intermediolateral nucleus in the lateral grey column of the spinal cord are carried in the white ramus communicans to the paravertebral ganglia of the sympathetic trunk. Once the preganglionic nerve has traversed a white ramus communicans, it can do one of three things. * The preganglionic neuron can synapse with a postganglionic sympathetic neuron in the sympathetic paravertebral ganglion at that level. From here, the postganglionic sympathetic neuron can travel back out the grey ramus communicans ...
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White Ramus Communicans
The white ramus communicans (plural: rami communicantes) from Latin ramus (branch) and communicans (communicating) is the preganglionic sympathetic outflow nerve tract from the spinal cord. Each of the thoracic, and the first and second lumbar nerves contribute a white ramus communicans to the adjoining sympathetic ganglion, unlike the gray rami which are located at each spinal level. White rami communicantes contain both myelinated and unmyelinated preganglionic sympathetic fibers, ( GVE and GVA). The white ramus appears white because there are more myelinated than unmyelinated fibers unlike the gray rami. Structure The white rami communicantes are the preganglionic sympathetic outflow from the spinal cord. The cell bodies for the preganglionic sympathetic myelinated fibers in the white rami communicantes lie in the ipsilateral (same sided) intermediolateral cell column in the spinal cord which extends from T1-L2. These rami also contain general visceral afferent fibers (sen ...
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Autonomic Nervous System
The autonomic nervous system (ANS), formerly referred to as the vegetative nervous system, is a division of the peripheral nervous system that supplies viscera, internal organs, smooth muscle and glands. The autonomic nervous system is a control system that acts largely unconsciously and regulates bodily functions, such as the heart rate, its force of contraction, digestion, respiratory rate, pupillary dilation, pupillary response, Micturition, urination, and sexual arousal. This system is the primary mechanism in control of the fight-or-flight response. The autonomic nervous system is regulated by integrated reflexes through the brainstem to the spinal cord and organ (anatomy), organs. Autonomic functions include control of respiration, heart rate, cardiac regulation (the cardiac control center), vasomotor activity (the vasomotor center), and certain reflex, reflex actions such as coughing, sneezing, swallowing and vomiting. Those are then subdivided into other areas and are also ...
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Sympathetic Nervous System
The sympathetic nervous system (SNS) is one of the three divisions of the autonomic nervous system, the others being the parasympathetic nervous system and the enteric nervous system. The enteric nervous system is sometimes considered part of the autonomic nervous system, and sometimes considered an independent system. The autonomic nervous system functions to regulate the body's unconscious actions. The sympathetic nervous system's primary process is to stimulate the body's fight or flight response. It is, however, constantly active at a basic level to maintain homeostasis. The sympathetic nervous system is described as being antagonistic to the parasympathetic nervous system which stimulates the body to "feed and breed" and to (then) "rest-and-digest". Structure There are two kinds of neurons involved in the transmission of any signal through the sympathetic system: pre-ganglionic and post-ganglionic. The shorter preganglionic neurons originate in the thoracolumbar division o ...
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Postganglionic Nerve Fibers
In the autonomic nervous system, fibers from the ganglion to the effector organ are called postganglionic fibers. Neurotransmitters The neurotransmitters of postganglionic fibers differ: * In the parasympathetic division, neurons are ''cholinergic''. That is to say acetylcholine is the primary neurotransmitter responsible for the communication between neurons on the parasympathetic pathway. * In the sympathetic division, neurons are mostly ''adrenergic'' (that is, epinephrine and norepinephrine function as the primary neurotransmitters). Notable exceptions to this rule include the sympathetic innervation of sweat glands and arrectores pilorum muscles where the neurotransmitter at both pre and post ganglionic synapses is acetylcholine. Another notable structure is the medulla of the adrenal gland, where chromaffin cells function as modified post-ganglionic nerves. Instead of releasing epinephrine and norepinephrine into a synaptic cleft, these cells of the adrenal medulla rel ...
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Intermediolateral Cell Column
The intermediolateral nucleus (IML) is a region of grey matter found in one of the three grey columns of the spinal cord, the lateral grey column. This is Rexed lamina VII. The intermediolateral cell column exists at vertebral levels T1 – L3. It mediates the entire sympathetic innervation of the body, but the nucleus resides in the grey matter of the spinal cord. Rexed Lamina VII contains several well defined nuclei including the nucleus dorsalis (Clarke's column), the intermediolateral nucleus, and the sacral autonomic nucleus. It extends from T1 to L3, and contains the autonomic motor neurons that give rise to the preganglionic fibers of the sympathetic nervous system, (preganglionic sympathetic general visceral efferent General visceral efferent fibers (GVE) or visceral efferents or autonomic efferents, are the efferent nerve fibers of the autonomic nervous system (also known as the ''visceral efferent nervous system'' that provide motor innervation to smooth m ...s). ...
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Preganglionic Nerve Fibers
In the autonomic nervous system, fibers from the CNS to the ganglion are known as preganglionic fibers. All preganglionic fibers, whether they are in the sympathetic division or in the parasympathetic division, are cholinergic (that is, these fibers use acetylcholine as their neurotransmitter) and they are myelinated. Sympathetic preganglionic fibers tend to be shorter than parasympathetic preganglionic fibers because sympathetic ganglia are often closer to the spinal cord than are the parasympathetic ganglia. Another major difference between the two ANS (autonomic nervous systems) is divergence. Whereas in the parasympathetic division there is a divergence factor of roughly 1:4, in the sympathetic division there can be a divergence of up to 1:20. This is due to the number of synapses formed by the preganglionic fibers with ganglionic neurons. See also * Postganglionic fibers In the autonomic nervous system, fibers from the ganglion to the effector organ are call ...
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