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Quagga Mussel
The quagga mussel (''Dreissena rostriformis'', also known as ''Dreissena bugensis'' or ''Dreissena rostriformis bugensis'') is a species (or subspecies) of freshwater mussel, an aquatic bivalve mollusk in the family Dreissenidae. It has an average lifespan of 3 to 5 years. The species is indigenous to the Dnieper River drainage of Ukraine, and is named after the quagga, an extinct subspecies of African zebra, possibly because, like the quagga, its stripes fade out towards the ventral side. The invasive quagga mussel is currently of major concern as it spreads in the rivers and lakes of Europe and also in the Great Lakes of North America where it was brought to by overseas shippers that use the Saint Lawrence Seaway. Appearance The quagga mussel shell is generally black, yellow, and/or zig-zagged. However, a large range of shell morphologies is seen, including a distinct morph in Lake Erie that is pale or completely white. The shell has a rounded carina and a convex ventral ...
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Zebra Mussel
The zebra mussel (''Dreissena polymorpha'') is a small freshwater mussel. The species originates from the lakes of southern Russia and Ukraine, but has been accidentally introduced to numerous other areas and has become an invasive species in many countries worldwide. Since the 1980s, the species has invaded the Great Lakes, Hudson River, and Lake Travis. The species was first described in 1769 by German zoologist Peter Simon Pallas in the Ural, Volga, and Dnieper Rivers. Zebra mussels get their name from a striped pattern commonly seen on their shells, though it is not universally present. They are usually about the size of a fingernail, but can grow to a maximum length around . Shells are D-shaped, and attached to the substrate with strong byssal fibers, which come out of their umbo on the dorsal (hinged) side. Ecology Zebra mussels and the closely related and ecologically similar quagga mussels are filter-feeding organisms; they remove particles from the water column. ...
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Ligament (bivalve)
A hinge ligament is a crucial part of the anatomical structure of a bivalve shell, i.e. the shell of a bivalve mollusk. The shell of a bivalve has two valves and these are joined together by the ligament at the dorsal edge of the shell. The ligament is made of a strong, flexible and elastic, fibrous, proteinaceous material which is usually pale brown, dark brown or black in color. In life, the shell needs to be able to open a little (to allow the foot and siphons to protrude) and then close again. As well as connecting the two bivalve shells together at the hinge line, the ligament also functions as a spring which automatically opens the valves when the adductor muscle or muscles (that close the valves) relax. Composition The ligament is an uncalcified elastic structure comprised in its most minimal state of two layers: a lamellar layer and a fibrous layer. The lamellar layer consists entirely of organic material (a protein and collagen matrix), is generally brown in color, and ...
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Danube River
The Danube ( ; ) is a river that was once a long-standing frontier of the Roman Empire and today connects 10 European countries, running through their territories or being a border. Originating in Germany, the Danube flows southeast for , passing through or bordering Austria, Slovakia, Hungary, Croatia, Serbia, Romania, Bulgaria, Moldova, and Ukraine before draining into the Black Sea. Its drainage basin extends into nine more countries. The largest cities on the river are Vienna, Budapest, Belgrade and Bratislava, all of which are the capitals of their respective countries; the Danube passes through four capital cities, more than any other river in the world. Five more capital cities lie in the Danube's basin: Bucharest, Sofia, Zagreb, Ljubljana and Sarajevo. The fourth-largest city in its basin is Munich, the capital of Bavaria, standing on the Isar River. The Danube is the second-longest river in Europe, after the Volga in Russia. It flows through much of Central and Sou ...
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United States Geological Survey
The United States Geological Survey (USGS), formerly simply known as the Geological Survey, is a scientific agency of the United States government. The scientists of the USGS study the landscape of the United States, its natural resources, and the natural hazards that threaten it. The organization's work spans the disciplines of biology, geography, geology, and hydrology. The USGS is a fact-finding research organization with no regulatory responsibility. The agency was founded on March 3, 1879. The USGS is a bureau of the United States Department of the Interior; it is that department's sole scientific agency. The USGS employs approximately 8,670 people and is headquartered in Reston, Virginia. The USGS also has major offices near Lakewood, Colorado, at the Denver Federal Center, and Menlo Park, California. The current motto of the USGS, in use since August 1997, is "science for a changing world". The agency's previous slogan, adopted on the occasion of its hundredt ...
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Byssal Thread
A byssus () is a bundle of filaments secreted by many species of bivalve mollusc that function to attach the mollusc to a solid surface. Species from several families of clams have a byssus, including pen shells (Pinnidae), true mussels (Mytilidae), and Dreissenidae. Filaments Byssus filaments are created by certain kinds of marine and freshwater bivalve mollusks, which use the byssus to attach themselves to rocks, substrates, or seabeds. In edible mussels, the inedible byssus is commonly known as the "beard", and is removed before cooking. Many species of mussels secrete byssus threads to anchor themselves to surfaces, with families including the Arcidae, Mytilidae, Anomiidae, Pinnidae, Pectinidae, Dreissenidae, and Unionidae. Mechanics The byssus, or byssal complex, is composed of multiple extracellular collagenous threads that are placed radially by the mussel from a central stem. Each thread is composed of three regions: a corrugated proximal region close to the mussel b ...
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Dioecious
Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is biparental reproduction. Dioecy has costs, since only about half the population directly produces offspring. It is one method for excluding self-fertilization and promoting allogamy (outcrossing), and thus tends to reduce the expression of recessive deleterious mutations present in a population. Plants have several other methods of preventing self-fertilization including, for example, dichogamy, herkogamy, and self-incompatibility. Dioecy is a dimorphic sexual system, alongside gynodioecy and androdioecy. In zoology In zoology, dioecious species may be opposed to hermaphroditic species, meaning that an individual is either male or female, in which case the synonym gonochory is more often used. Most animal species are dioecious (gon ...
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Pseudofeces
Pseudofeces or pseudofaeces are a specialized method of expulsion that filter-feeding bivalve mollusks (and filter-feeding gastropod mollusks) use in order to get rid of suspended particles such as particles of grit which cannot be used as food, and which have been rejected by the animal. The rejected particles are wrapped in mucus, and are then expelled without having passed through the digestive tract.Shimek, Ronald L''Phytoplankton, A Necessity For Clams''/ref> Thus, although they may closely resemble the mollusk's real feces, they are not actually feces, hence the name pseudofeces, meaning false feces. Occurrence Bivalves which exhibit this behavior are numerous and include Ostreidae oysters (such as ''Crassostrea'') and Dreissenidae false mussels (such as ''Dreissena''). Gastropods which filter feed are in a minority, but include the mudsnail genus
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Mucus
Mucus ( ) is a slippery aqueous secretion produced by, and covering, mucous membranes. It is typically produced from cells found in mucous glands, although it may also originate from mixed glands, which contain both serous and mucous cells. It is a viscous colloid containing inorganic salts, antimicrobial enzymes (such as lysozymes), immunoglobulins (especially IgA), and glycoproteins such as lactoferrin and mucins, which are produced by goblet cells in the mucous membranes and submucosal glands. Mucus serves to protect epithelial cells in the linings of the respiratory, digestive, and urogenital systems, and structures in the visual and auditory systems from pathogenic fungi, bacteria and viruses. Most of the mucus in the body is produced in the gastrointestinal tract. Amphibians, fish, snails, slugs, and some other invertebrates also produce external mucus from their epidermis as protection against pathogens, and to help in movement and is also produced in fish to line the ...
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Veliger
A veliger is the planktonic larva of many kinds of sea snails and freshwater snails, as well as most bivalve molluscs (clams) and tusk shells. Description The veliger is the characteristic larva of the gastropod, bivalve and scaphopod taxonomic classes. It is produced following either the embryonic or trochophore larval stage of development. In bivalves the veliger is sometimes referred to as a D-stage (early in its development) or pediveliger (late in its development) larva. This stage in the life history of these groups is a free-living planktonic organism; this mode of life potentially enhances dispersal to new regions far removed from the adult mollusks that produced the larvae. The general structure of the veliger includes a shell that surrounds the visceral organs of the larva (e.g., digestive tract, much of the nervous system, excretory organs) and a ciliated velum that extends beyond the shell as a single or multi-lobed structure used for swimming and particula ...
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Algae
Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular microalgae, such as ''Chlorella,'' ''Prototheca'' and the diatoms, to multicellular forms, such as the giant kelp, a large brown alga which may grow up to in length. Most are aquatic and autotrophic (they generate food internally) and lack many of the distinct cell and tissue types, such as stomata, xylem and phloem that are found in land plants. The largest and most complex marine algae are called seaweeds, while the most complex freshwater forms are the ''Charophyta'', a division of green algae which includes, for example, ''Spirogyra'' and stoneworts. No definition of algae is generally accepted. One definition is that algae "have chlorophyll ''a'' as their primary photosynthetic pigment and lack a sterile covering of cells around thei ...
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Zooplankton
Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by currents in the ocean, or by currents in seas, lakes or rivers. Zooplankton can be contrasted with phytoplankton, which are the plant component of the plankton community ("phyto" comes from the Greek word for ''plant''). Zooplankton are heterotrophic (other-feeding), whereas phytoplankton are autotrophic (self-feeding). This means zooplankton cannot manufacture their own food but must eat other plants or animals instead — in particular they eat phytoplankton. Zooplankton are generally larger than phytoplankton, most are microscopic, but some (such as jellyfish) are macroscopic and can be seen with the naked eye. Many protozoans (single-celled protists that prey on other microscopic life) are zooplankton, including zooflagellates, fo ...
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Phytoplankton
Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Phytoplankton obtain their energy through photosynthesis, as do trees and other plants on land. This means phytoplankton must have light from the sun, so they live in the well-lit surface layers (euphotic zone) of oceans and lakes. In comparison with terrestrial plants, phytoplankton are distributed over a larger surface area, are exposed to less seasonal variation and have markedly faster turnover rates than trees (days versus decades). As a result, phytoplankton respond rapidly on a global scale to climate variations. Phytoplankton form the base of marine and freshwater food webs and are key players in the global carbon cycle. They account for about half of global photosynthetic activity and at least half of the oxygen production, despite ...
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