Protochordate
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Protochordate
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cephalochordates. These CSIs provi ...
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Endostyle
The endostyle is an anatomical feature found in invertebrate chordates and larval lampreys. It is an organ which assists chordates in filter-feeding. It is found in adult urochordates and cephalochordates, as well as in the larvae of the vertebrate lampreys, but has been transformed through evolutionary time into the thyroid in all other vertebrates. Since the endostyle is found in all three branches of chordates, it is presumed to have arisen in the common ancestor of these taxa, along with a shift to internal feeding for extracting suspended food from the water. The endostyle is found in the pharynx. The food particles suspended in the water adhere to the mucus it produces. The filtered water is expelled through the gill slits, while the food and mucus is then passed, by the sweeping movement of cilia that coats the endostyle, through the pharynx of the organism and into the esophagus. The endostyle in larval lampreys (ammocoetes) metamorphoses into the thyroid gland in adults ...
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Tunicate
A tunicate is a marine invertebrate animal, a member of the subphylum Tunicata (). It is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates). The subphylum was at one time called Urochordata, and the term urochordates is still sometimes used for these animals. They are the only chordates that have lost their myomeric segmentation, with the possible exception of the 'seriation of the gill slits'. Some tunicates live as solitary individuals, but others replicate by budding and become colonies, each unit being known as a zooid. They are marine filter feeders with a water-filled, sac-like body structure and two tubular openings, known as siphons, through which they draw in and expel water. During their respiration and feeding, they take in water through the incurrent (or inhalant) siphon and expel the filtered water through the excurrent (or exhalant) siphon. Most adult tunicates are sessile, immobile and perman ...
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Pikaiidae
''Pikaia gracilens'' is an extinct, primitive chordate animal known from the Middle Cambrian Burgess Shale of British Columbia. Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became the "most famous early chordate fossil," or "famously known as the earliest" chordates. It is estimated to have lived during the latter period of the Cambrian explosion. Since it initial discovery, more than a hundred specimens have been recovered. The body structure resembles that of the lancelet and perhaps it swam much like an eel. A notochord and myomeres (segmented blocks of skeletal muscles) span the entire length of the body, and are considered as the defining signatures of chordate characters. Its primitive feature is indicated by the body covering, a cuticle, which is characteristic of invertebrates and some protochordates. The exact phylogenetic position is unclear. Proposed affinities include th ...
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Cambrian Stage 3
Cambrian Stage 3 is the still unnamed third stage of the Cambrian. It succeeds Cambrian Stage 2 and precedes Cambrian Stage 4, although neither its base nor top have been formally defined. The plan is for its lower boundary to correspond approximately to the first appearance of trilobites, about million years ago, though the globally asynchronous appearance of trilobites warrants the use of a separate, globally synchronous marker to define the base. The upper boundary and beginning of Cambrian Stage 4 is informally defined as the first appearance of the trilobite genera ''Olenellus'' or '' Redlichia'' around million years ago. Naming The International Commission on Stratigraphy has not officially named the 3rd stage of the Cambrian. The stage approximately corresponds to the "Atdabanian", which is used by geologists working in Siberia. Biostratigraphy The oldest trilobite known is ''Lemdadella'' which appears at the beginning of the '' Fallotaspis'' zone. The Cambrian radiati ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Conserved Signature Indels
Conserved signature inserts and deletions (CSIs) in protein sequences provide an important category of molecular markers for understanding phylogenetic relationships. CSIs, brought about by rare genetic changes, provide useful phylogenetic markers that are generally of defined size and they are flanked on both sides by conserved regions to ensure their reliability. While indels can be arbitrary inserts or deletions, CSIs are defined as only those protein indels that are present within conserved regions of the protein. The CSIs that are restricted to a particular clade or group of species, generally provide good phylogenetic markers of common evolutionary descent. Due to the rarity and highly specific nature of such changes, it is less likely that they could arise independently by either convergent or parallel evolution (i.e. homoplasy) and therefore are likely to represent synapomorphy. Other confounding factors such as differences in evolutionary rates at different sites or among ...
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Metameric
In biology, metamerism is the phenomenon of having a linear series of body segments fundamentally similar in structure, though not all such structures are entirely alike in any single life form because some of them perform special functions. In animals, metameric segments are referred to as somites or metameres. In plants, they are referred to as metamers or, more concretely, phytomers. In animals In animals, zoologists define metamery as a mesodermal event resulting in serial repetition of unit subdivisions of ectoderm and mesoderm products. Endoderm is not involved in metamery. Segmentation is not the same concept as metamerism: segmentation can be confined only to ectodermally derived tissue, e.g., in the Cestoda tapeworms. Metamerism is far more important biologically since it results in metameres - also called somites - that play a critical role in advanced locomotion. One can divide metamerism into two main categories: * homonomous metamery is a strict serial successio ...
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Circulatory System
The blood circulatory system is a system of organs that includes the heart, blood vessels, and blood which is circulated throughout the entire body of a human or other vertebrate. It includes the cardiovascular system, or vascular system, that consists of the heart and blood vessels (from Greek ''kardia'' meaning ''heart'', and from Latin ''vascula'' meaning ''vessels''). The circulatory system has two divisions, a systemic circulation or circuit, and a pulmonary circulation or circuit. Some sources use the terms ''cardiovascular system'' and ''vascular system'' interchangeably with the ''circulatory system''. The network of blood vessels are the great vessels of the heart including large elastic arteries, and large veins; other arteries, smaller arterioles, capillaries that join with venules (small veins), and other veins. The Closed circulatory system, circulatory system is closed in vertebrates, which means that the blood never leaves the network of blood vessels. Some in ...
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Coelom
The coelom (or celom) is the main body cavity in most animals and is positioned inside the body to surround and contain the digestive tract and other organs. In some animals, it is lined with mesothelium. In other animals, such as molluscs, it remains undifferentiated. In the past, and for practical purposes, coelom characteristics have been used to classify bilaterian animal phyla into informal groups. Etymology The term ''coelom'' derives from the Ancient Greek word (), meaning 'cavity'. Structure Development The coelom is the mesodermally lined cavity between the gut and the outer body wall. During the development of the embryo, coelom formation begins in the gastrulation stage. The developing digestive tube of an embryo forms as a blind pouch called the archenteron. In Protostomes, the coelom forms by a process known as schizocoely. The archenteron initially forms, and the mesoderm splits into two layers: the first attaches to the body wall or ectoderm, forming ...
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Bilateral Symmetry
Symmetry in biology refers to the symmetry observed in organisms, including plants, animals, fungi, and bacteria. External symmetry can be easily seen by just looking at an organism. For example, take the face of a human being which has a plane of symmetry down its centre, or a pine cone with a clear symmetrical spiral pattern. Internal features can also show symmetry, for example the tubes in the human body (responsible for transporting gases, nutrients, and waste products) which are cylindrical and have several planes of symmetry. Biological symmetry can be thought of as a balanced distribution of duplicate body parts or shapes within the body of an organism. Importantly, unlike in mathematics, symmetry in biology is always approximate. For example, plant leaves – while considered symmetrical – rarely match up exactly when folded in half. Symmetry is one class of patterns in nature whereby there is near-repetition of the pattern element, either by reflection or rotation ...
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Pharyngeal Slit
Pharyngeal slits are filter-feeding organs found among deuterostomes. Pharyngeal slits are repeated openings that appear along the pharynx caudal to the mouth. With this position, they allow for the movement of water in the mouth and out the pharyngeal slits. It is postulated that this is how pharyngeal slits first assisted in filter-feeding, and later, with the addition of gills along their walls, aided in respiration of aquatic chordates. These repeated segments are controlled by similar developmental mechanisms. Some hemichordate species can have as many as 200 gill slits. Pharyngeal clefts resembling gill slits are transiently present during the embryonic stages of tetrapod development. The presence of pharyngeal arches and clefts in the neck of the developing human embryo famously led Ernst Haeckel to postulate that "ontogeny recapitulates phylogeny"; this hypothesis, while false, contains elements of truth, as explored by Stephen Jay Gould in ''Ontogeny and Phylogeny''.. A ...
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Thyroid
The thyroid, or thyroid gland, is an endocrine gland in vertebrates. In humans it is in the neck and consists of two connected lobes. The lower two thirds of the lobes are connected by a thin band of tissue called the thyroid isthmus. The thyroid is located at the front of the neck, below the Adam's apple. Microscopically, the functional unit of the thyroid gland is the spherical thyroid follicle, lined with follicular cells (thyrocytes), and occasional parafollicular cells that surround a lumen containing colloid. The thyroid gland secretes three hormones: the two thyroid hormones triiodothyronine (T3) and thyroxine (T4)and a peptide hormone, calcitonin. The thyroid hormones influence the metabolic rate and protein synthesis, and in children, growth and development. Calcitonin plays a role in calcium homeostasis. Secretion of the two thyroid hormones is regulated by thyroid-stimulating hormone (TSH), which is secreted from the anterior pituitary gland. TSH is regula ...
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