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Ophioglossaceae
Ophioglossaceae, the adder's-tongue family, is a small family of ferns. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only family in the order Ophioglossales, which together with the Psilotales is placed in the subclass Ophioglossidae. The Ophioglossidae are one of the groups traditionally known as eusporangiate ferns. Members of the family differ from other ferns in a number of ways. Many have only a single fleshy leaf at a time. Their gametophytes are subterranean and rely on fungi for energy. Description Members of Ophioglossaceae are usually terrestrial (excepting a few epiphytic species of ''Ophioglossum'') and occur in both temperate and tropical areas. They differ from the other ferns in several respects: * Many species only send up one frond or leaf-blade per year, producing only a single leaf at a time. The leaves are usually fleshy, and in temperate areas will often turn brownish or reddish during colder months. * Instead of the leptosp ...
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Eusporangiate Fern
Eusporangiate ferns are vascular spore plants, whose sporangia arise from several epidermal cells and not from a single cell as in leptosporangiate ferns. Typically these ferns have reduced root systems and sporangia that produce large amounts of spores (up to 7000 spores per sporangium in '' Christensenia'') There are four extant eusporangiate fern families, distributed among three classes. Each family is assigned to its own order. *Class Psilotopsida **Order Psilotales, family Psilotaceae – Whisk ferns (2 genera, about 17 species) **Order Ophioglossales, family Ophioglossaceae – Adder's-tongues (5 genera, about 80 species) *Class Equisetopsida **Order Equisetales, family Equisetaceae – Horsetails (1 genus, about 15 species) *Class Marattiopsida **Order Marattiales, family Marattiaceae – Marattoid ferns (6 genera, about 500 species) The following diagram shows a likely phylogenic placement of eusporangiate fern classes within the vascular plants. Cladistics While it ...
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Botrychium Lunaria Kz01
''Botrychium'' is a genus of ferns, seedless vascular plants in the family Ophioglossaceae. ''Botrychium'' species are known as moonworts. They are small, with fleshy roots, and reproduce by spores shed into the air. One part of the leaf, the trophophore, is sterile and fernlike; the other, the sporophore, is fertile and carries the clusters of sporangia or spore cases. Some species only occasionally emerge above ground and gain most of their nourishment from an association with mycorrhizal fungi. The circumscription of ''Botrychium'' is disputed between different authors; some botanists include the genera '' Botrypus'' and '' Sceptridium'' within ''Botrychium'', while others treat them as distinct. The latter treatment is provisionally followed here. Taxonomy * – pointed moonwort * – Alaska moonwort * – upswept moonwort, triangle-lobed moonwort, upward-lobed moonwort * – northern moonwort * – prairie moonwort, prairie dunewort, Iowa moonwort * – dainty moo ...
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Moonwort
''Botrychium'' is a genus of ferns, seedless vascular plants in the family Ophioglossaceae. ''Botrychium'' species are known as moonworts. They are small, with fleshy roots, and reproduce by spores shed into the air. One part of the leaf, the trophophore, is sterile and fernlike; the other, the sporophore, is fertile and carries the clusters of sporangia or spore cases. Some species only occasionally emerge above ground and gain most of their nourishment from an association with mycorrhizal fungi. The circumscription of ''Botrychium'' is disputed between different authors; some botanists include the genera '' Botrypus'' and '' Sceptridium'' within ''Botrychium'', while others treat them as distinct. The latter treatment is provisionally followed here. Taxonomy * – pointed moonwort * – Alaska moonwort * – upswept moonwort, triangle-lobed moonwort, upward-lobed moonwort * – northern moonwort * – prairie moonwort, prairie dunewort, Iowa moonwort * – dainty moo ...
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Ophioglossum
''Ophioglossum'', the adder's-tongue ferns, is a genus of about 50 species of ferns A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except th ... in the family Ophioglossaceae. The name ''Ophioglossum'' comes from the Greek language, Greek meaning "snake-tongue".eFloras: ''Ophioglossum''
accessed 14 February 2014.
Their cosmopolitan distribution is mainly in tropical and subtropical habitats. The genus has the largest number of chromosomes in the known plant kingdom, but contrary to popular belief does not have the largest number of chromosomes out of all known organisms, falling short to the pr ...
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Ophioglossum Vulgatum
''Ophioglossum vulgatum'', commonly known as adder's-tongue, southern adders-tongue or adders-tongue fern, is a species of fern in the family ''Ophioglossaceae''. The adder’s tongue fern is generally believed to have the largest number of chromosomes wit1262 compared to the human’s 46. Distribution It is native to many regions with a wide scattered distribution: throughout temperate through tropical Africa and throughout the temperate Northern Hemisphere in Europe, northeastern North America, temperate Asia, and Eurasia. This small, hard-to-spot plant can occur singly in unimproved pastures, rock crevices and grassy path-sides, but also can occur in colonies of hundreds of plants in sand dunes. Description ''Ophioglossum vulgatum'' grows from a rhizome base to 10–20 cm tall (rarely to 30 cm). It consists of a two-part frond, separated into a rounded diamond-shaped sheath and narrow spore-bearing spike. The spike has around 10-40 segments on each side. It repro ...
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Grape Fern
''Sceptridium'' is a genus of seedless vascular plants in the family Ophioglossaceae, closely allied to (and often included as a subgenus of) the genus '' Botrychium'' (the moonworts and grapeferns). It is also closely related to the genus '' Botrypus'' (the rattlesnake fern, often treated as the subgenus ''Osmundopteris'' under ''Botrychium''). ''Sceptridium'' species are commonly called the grape-ferns. These plants are small with fleshy roots, and reproduce by spores shed into the air. They differ from the moonworts in having at least some sterile fronds (all fronds in ''Botrychium'' are spore-bearing), and in the fronds being bi- or tri-pinnate (''Botrychium'' are single pinnate, or rarely bipinnate); and from ''Botrypus'' in being evergreen, or at least winter-green (''Botrypus'' are deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally s ...
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Ophioglossum Malviae
''Ophioglossum malviae'' is a species of fern in the genus ''Ophioglossum'' and found in the forests of Ahwa in the Dang District, India, Dang District of Gujarat. It is claimed to be the world's smallest land fern, with the entire plant growing to a size of about only 1 to 1.5 cm. References

Ophioglossaceae {{fern-stub ...
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Ophioglossum Reticulatum
''Ophioglossum reticulatum'', the netted adder's-tongue, is a species of fern in the family Ophioglossaceae. It has a pantropical/pansubtropical distribution; Latin America, the Caribbean, Sub-Saharan Africa, Madagascar, Yemen, the Indian Subcontinent, Southeast Asia, warmer parts of China, Malesia, Korea, Japan, and many tropical islands. A hexaploid Polyploidy is a condition in which the cells of an organism have more than one pair of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contains ..., it has the highest number of chromosomes of any plant, 720. Its leaves—or leaf, individuals only grow one per year—are edible, and are regularly consumed by people in Africa and Asia. References reticulatum Ferns of the Americas Ferns of Africa Ferns of Asia Ferns of Oceania Flora of Mexico Flora of Central America Flora of the Caribbean Flora of South America Flora ...
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Helminthostachys
''Helminthostachys'' is a fern genus in the Ophioglossaceae (Adder's tongue) family.details
Tropicos.org. Missouri Botanical Garden. 15 Jan 2012 It is widespread throughout southeast and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
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References

Ophioglossaceae
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Fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Gametophyte
A gametophyte () is one of the two alternation of generations, alternating multicellular organism, multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the Sexual reproduction of plants, sexual phase in the life cycle of plants and algae. It develops sex organs that produce gametes, haploid sex cells that participate in fertilization to form a diploid zygote which has a double set of chromosomes. Cell division of the zygote results in a new diploid multicellular organism, the second stage in the life cycle known as the sporophyte. The sporophyte can produce haploid spores by meiosis that on germination produce a new generation of gametophytes. Algae In some multicellular green algae (''Ulva lactuca'' is one example), red algae and brown algae, sporophytes and gametophytes may be externally indistinguishable (isomorphic). In ''Ulva (genus), Ulv ...
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Pteridophyte Phylogeny Group
The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relationships discovered through phylogenetic studies. In 2016, the group published a classification for extant pteridophytes, termed "PPG I". The paper had 94 authors (26 principal and 68 additional). PPG I A first classification, PPG I, was produced in 2016, covering only extant (living) pteridophytes. The classification was rank-based, using the ranks of class, subclass, order, suborder, family, subfamily and genus. Phylogeny The classification was based on a consensus phylogeny, shown below to the level of order. The very large order Polypodiales was divided into two suborders, as well as families not placed in a suborder: Classification to subfamily level To the level of subfamily, the PPG I classification is as follows. *Class Lycopodi ...
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