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Nematomorpha
Nematomorpha (sometimes called Gordiacea, and commonly known as horsehair worms, hairsnakes, or Gordian worms) are a phylum of parasitoid animals superficially similar to nematode worms in morphology, hence the name. Most species range in size from long, reaching 2 metres in extreme cases, and in diameter. Horsehair worms can be discovered in damp areas, such as watering troughs, swimming pools, streams, puddles, and cisterns. The adult worms are free-living, but the larvae are parasitic on arthropods, such as beetles, cockroaches, mantises, orthopterans, and crustaceans. About 351 freshwater species are known and a conservative estimate suggests that there may be about 2000 freshwater species worldwide. The name "Gordian" stems from the legendary Gordian knot. This relates to the fact that nematomorphs often coil themselves in tight balls that resemble knots. Description and biology Nematomorphs possess an external cuticle without cilia. Internally, they have only longitudina ...
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Nectonematoidea
''Nectonema'' is a genus of marine horsehair worms first described by Addison E. Verrill in 1879. It is the only genus in the family Nectonematidae described by Henry B. Ward in 1892, in the order Nectonematoidea, and in the class Nectonematoida. The genus contains five species; all species have a parasitic larval stage inhabiting crustacean hosts and a free-living adult stage that swims in open water. Taxonomy Nectonematoidea is one of two orders within the phylum Nematomorpha, the other being Gordioidea. The latter is likewise in a single class Gordioida, which is a significantly larger taxon, with over 300 known species. Nematomorpha are known as horsehair worms or Gordian worms, and form a sister-group to the nematodes. The following classification shows the place of Nectonematoida within the protostomes according to Minelli (2008) and Tedersoo (2017): Within Nectonematoida only a single genus, ''Nectonema'', is known, with five species so far described: * '' Nectone ...
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Nematode
The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant- parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broad range of environments. Less formally, they are categorized as Helminths, but are taxonomically classified along with arthropods, tardigrades and other moulting animals in the clade Ecdysozoa, and unlike flatworms, have tubular digestive systems with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum. Nematode species can be difficult to distinguish from one another. Consequently, estimates of the number of nematode species described to date vary by author and may change rapidly over time. A 2013 survey of animal biodiversity published in the mega ...
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Paragordius Tricuspidatus
''Paragordius tricuspidatus'' belongs to the phylum Nematomorpha, and is known for manipulating the behavior of its host, the cricket ''Nemobius sylvestris''. In its larval stage, the worm is microscopic, but grows into a large worm () inside its host after accidental ingestion since their eggs are laid at the edge of the water by rivers where crickets frequently reside. Upon ingestion, the worm nourishes upon its host and fills the entire body cavity of the cricket, until maturation, when the parasitic worm is ready to exit into water to complete its life cycle, maximizing its reproductive success. The worm induces a peculiar behavior on its cricket host, which causes it to leap into water whereby the parasitic worm can slither out and find its mate, while the cricket often perishes. Should the cricket be preyed upon by a predator, such as a fish or frog, the worm has the ability to not only escape from the host's body but also the predator's digestive system. The worm emerges f ...
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Gordioidea
Gordioidea is an order (sometimes placed at superfamily level) of parasitic horsehair worms. Its taxonomy remains uncertain, but appears to be contained in the monotypic class Gordioida and contains about 320 known species. Biology Gordioidean adults are free-living in freshwater or semi-terrestrial habitats and larvae parasitise insects, primarily Orthopterans. Unlike Nectonematoideans, which are marine, gordioideans lack lateral rows of setae and have a single, ventral epidermal cord and their blastocoel is filled with mesenchyme in young worms but become open when older.Pechenik (2010) ''Biology of the Invertebrates'', p. 457. Families and genera The Integrated Taxonomic Information System lists the following genera in two families: Chordodidae Auth. May, 1919; selected genera: ;Subfamily Chordodinae Heinze, 1935 * '' Chordodes'' Creplin, 1847 * '' Dacochordodes'' Capuse, 1965 * ''Euchordodes'' Heinze, 1937 * ''Neochordodes'' Carvalho, 1942 * ''Pantachordodes'' Heinze, 195 ...
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Phylum
In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained un ...
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Parasitoid
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable strategy, evolutionary strategies within parasitism, distinguished by the fatal prognosis for the host, which makes the strategy close to predation. Among parasitoids, strategies range from living inside the host (''endoparasitism''), allowing it to continue growing before emerging as an adult, to Paralysis, paralysing the host and living outside it (''ectoparasitism''). Hosts can include other parasitoids, resulting in hyperparasitism; in the case of oak galls, up to five levels of parasitism are possible. Some parasitoids Behavior-altering parasite, influence their host's behaviour in ways that favour the propagation of the parasitoid. Parasitoids are found in a variety of Taxon, taxa across the insect superorder Endopterygota, whose compl ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Larva
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The larva's appearance is generally very different from the adult form (''e.g.'' caterpillars and butterflies) including different unique structures and organs that do not occur in the adult form. Their diet may also be considerably different. Larvae are frequently adapted to different environments than adults. For example, some larvae such as tadpoles live almost exclusively in aquatic environments, but can live outside water as adult frogs. By living in a distinct environment, larvae may be given shelter from predators and reduce competition for resources with the adult population. Animals in the larval stage will consume food to fuel their transition into the adult form. In some organisms like polychaetes and barnacles, adults are ...
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Parasite
Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has characterised parasites as "predators that eat prey in units of less than one". Parasites include single-celled protozoans such as the agents of malaria, sleeping sickness, and amoebic dysentery; animals such as hookworms, lice, mosquitoes, and vampire bats; fungi such as honey fungus and the agents of ringworm; and plants such as mistletoe, dodder, and the broomrapes. There are six major parasitic strategies of exploitation of animal hosts, namely parasitic castration, directly transmitted parasitism (by contact), trophicallytransmitted parasitism (by being eaten), vector-transmitted parasitism, parasitoidism, and micropredation. One major axis of classification concerns invasiveness: an endoparasite lives inside the host's body ...
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Nerve Ring
A circumesophageal or circumpharyngeal nerve ring is an arrangement of nerve ganglia around the esophagus/ pharynx of an animal. It is a common feature of nematodes, molluscs, and many other invertebrate animals, though it is absent in all vertebrate animals and is not structurally possible in simpler ones such as water bears. The nerve ring, also called a nerve collar, creates a complete and closed loop around the food-entry parts of the animal's anatomy. In a typical molluscan arrangement, these include the cerebral, pedal, and pleural ganglia, with the esophagus passing through the center of the ring. In nematodes, the ring consists of only two to four large associative cells connected to two paired lateral ganglia, two ventral ganglia, and a single unpaired dorsal ganglion. Among arthropods, the usual arrangement is a single ganglion (the cerebral) positioned above the esophagus, a single ganglion or nerve mass (the subesophageal) below it, and commissures connecting the two ...
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Excretory System
The excretory system is a passive biological system that removes excess, unnecessary materials from the body fluids of an organism, so as to help maintain internal chemical homeostasis and prevent damage to the body. The dual function of excretory systems is the elimination of the waste products of metabolism and to drain the body of used up and broken down components in a liquid and gaseous state. In humans and other amniotes (mammals, birds and reptiles) most of these substances leave the body as urine and to some degree exhalation, mammals also expel them through sweating. Only the organs specifically used for the excretion are considered a part of the excretory system. In the narrow sense, the term refers to the urinary system. However, as excretion involves several functions that are only superficially related, it is not usually used in more formal classifications of anatomy or function. As most healthy functioning organs produce metabolic and other wastes, the entire org ...
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Respiratory System
The respiratory system (also respiratory apparatus, ventilatory system) is a biological system consisting of specific organs and structures used for gas exchange in animals and plants. The anatomy and physiology that make this happen varies greatly, depending on the size of the organism, the environment in which it lives and its evolutionary history. In land animals the respiratory surface is internalized as linings of the lungs. Gas exchange in the lungs occurs in millions of small air sacs; in mammals and reptiles these are called alveoli, and in birds they are known as atria. These microscopic air sacs have a very rich blood supply, thus bringing the air into close contact with the blood. These air sacs communicate with the external environment via a system of airways, or hollow tubes, of which the largest is the trachea, which branches in the middle of the chest into the two main bronchi. These enter the lungs where they branch into progressively narrower secondary ...
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