Myriapod
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Myriapod
Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial. The fossil record of myriapods reaches back into the late Silurian, although molecular evidence suggests a diversification in the Cambrian Period, and Cambrian fossils exist which resemble myriapods. The oldest unequivocal myriapod fossil is of the millipede ''Pneumodesmus newmani'', from the late Silurian (428 million years ago). ''P. newmani'' is also important as the earliest known terrestrial animal. The phylogenetic classification of myriapods is still debated. The scientific study of myriapods is myriapodology, and those who study myriapods are myriapodologists. Anatomy Myriapods have a single pair of antennae and, in most cases, simple eyes. Exceptions are the two classes symphylans and pauropods, and the millipede order Polydesmida and the centipede order Geophilomorpha, which are all eyele ...
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Millipede
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from the Latin for "thousand feet", no species was known to have 1,000 or more until the discovery of ''Eumillipes persephone'', which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures. Most millipedes are slow-moving detritivores, eating decaying leaves and other dead plant matter. Some eat fungi or ...
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Millipede
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from the Latin for "thousand feet", no species was known to have 1,000 or more until the discovery of ''Eumillipes persephone'', which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures. Most millipedes are slow-moving detritivores, eating decaying leaves and other dead plant matter. Some eat fungi or ...
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Diplopoda
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from the Latin for "thousand feet", no species was known to have 1,000 or more until the discovery of '' Eumillipes persephone'', which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures. Most millipedes are slow-moving detritivores, eating decaying leaves and other dead plant matter. Some eat fungi o ...
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Symphyla
Symphylans, also known as garden centipedes or pseudocentipedes, are soil-dwelling arthropods of the class Symphyla in the subphylum Myriapoda. Symphylans resemble centipedes, but are very small, non-venomous, and only distantly related to both centipedes and millipedes. They can move rapidly through the pores between soil particles, and are typically found from the surface down to a depth of about . They consume decaying vegetation, but can do considerable harm in an agricultural setting by consuming seeds, roots, and root hairs in cultivated soil. Juveniles have six pairs of legs, but over a lifetime of several years, they add an additional pair at each moult so an adult instar usually has twelve pairs of legs. Most adult symphylans have twelve leg pairs, but the first pair is absent or vestigial in some species (e.g., those in the genus ''Symphylella''), so some adults have only eleven leg pairs instead. Symphylans lack eyes. Their long antennae serve as sense organs. They h ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Myriapodology
Myriapodology is the scientific study of myriapods which includes centipedes and millipedes. The field of myriapodology can also cover other myriapods such as pauropods and symphylans. Those who study myriapods are myriapodologists. Societies * International Society of Myriapodology Journals * ''International Journal of Myriapodology'' * ''Myriapodologica'' * ''Myriapod Memoranda'' Notable myriapodologists * Carl Attems (1868–1952), Austrian zoologist, described over 1,000 species * Stanley Graham Brade-Birks (1887-1982), English myriapodologist who with Hilda K Brade-Birks authored ''Notes on Myriapoda'': 23 papers jointly from 1916 to the 1920s; then twelve more solo until 1939 * Henry W. Brolemann (1860–1933), French myriapodologist, described around 500 species * Ralph Vary Chamberlin (1879–1967), American arachnologist and myriapodologist, described over 1,000 species * Orator F. Cook (1867–1949), American botanist and myriapodologist, co-described world's leggiest ...
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Pauropoda
Pauropods are small, pale, millipede-like arthropods. Around 830 species in twelve families are found worldwide, living in soil and leaf mold. They look rather like centipedes, or millipedes, and may be a sister group of the latter. However, this is controversial, as a close relationship with Symphyla has also been posited. Anatomy and ecology Pauropods are soft, cylindrical animals with bodies long. They have neither eyes nor hearts, although they do have sensory organs which can detect light. The body segments have ventral tracheal/spiracular pouches forming apodemes similar to those in millipedes and Symphyla, although the trachea usually connected to these structures are absent in most species. There are long sensory hairs located throughout the body segments. Pauropods can usually be identified because of their distinctive anal plate, which is unique to pauropods. Different species of pauropods can be identified based on the size and shape of their anal plate. The antenn ...
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Chilopoda
Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (metameric) creatures with one pair of legs per body segment. All centipedes are venomous and can inflict painful bites, injecting their venom through pincer-like appendages known as forcipules. Despite the name, centipedes can have a varying number of legs, ranging from 30 to 382. Centipedes always have an odd number of pairs of legs; no centipede has exactly 100. Like spiders and scorpions, centipedes are predominantly carnivorous. Their size ranges from a few millimetres in the smaller lithobiomorphs and geophilomorphs to about in the largest scolopendromorphs. Centipedes can be found in a wide variety of environments. They ...
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Centipede
Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (metameric) creatures with one pair of legs per body segment. All centipedes are venomous and can inflict painful bites, injecting their venom through pincer-like appendages known as forcipules. Despite the name, centipedes can have a varying number of legs, ranging from 30 to 382. Centipedes always have an odd number of pairs of legs; no centipede has exactly 100. Like spiders and scorpions, centipedes are predominantly carnivorous. Their size ranges from a few millimetres in the smaller lithobiomorphs and geophilomorphs to about in the largest scolopendromorphs. Centipedes can be found in a wide variety of environments. They ...
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Centipede
Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (metameric) creatures with one pair of legs per body segment. All centipedes are venomous and can inflict painful bites, injecting their venom through pincer-like appendages known as forcipules. Despite the name, centipedes can have a varying number of legs, ranging from 30 to 382. Centipedes always have an odd number of pairs of legs; no centipede has exactly 100. Like spiders and scorpions, centipedes are predominantly carnivorous. Their size ranges from a few millimetres in the smaller lithobiomorphs and geophilomorphs to about in the largest scolopendromorphs. Centipedes can be found in a wide variety of environments. They ...
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Pneumodesmus Newmani
''Pneumodesmus newmani'' is a species of myriapod that lived in the Paleozoic. Its exact age is uncertain; it was originally interpreted as living , in the Late Silurian; however, subsequent research dates it to around 414 million years old, in the Early Devonian (Lochkovian). It is one of the first myriapods, and among the oldest creatures to have lived on land. It was discovered in 2004, and is known from a single specimen from Stonehaven, Aberdeenshire, Scotland. Discovery and naming The fossil of ''P. newmani'' was found by Mike Newman, a bus driver and amateur palaeontologist from Aberdeen, in a layer of sandstone rocks on the foreshore of Cowie, near Stonehaven. The species was later given the specific epithet "''newmani''" in honour of Newman. The holotype is kept in National Museum of Scotland, Edinburgh. The genus name is said to derived from the Greek ''pneumato'', meaning "air" or "breath", in reference to the inferred air-breathing habit. The proper word in ancient G ...
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Arthropod Mouthparts
The mouthparts of arthropods have evolved into a number of forms, each adapted to a different style or mode of feeding. Most mouthparts represent modified, paired appendages, which in ancestral forms would have appeared more like legs than mouthparts. In general, arthropods have mouthparts for cutting, chewing, piercing, sucking, shredding, siphoning, and filtering. This article outlines the basic elements of four arthropod groups: insects, myriapods, crustaceans and chelicerates. Insects are used as the model, with the novel mouthparts of the other groups introduced in turn. Insects are not, however, the ancestral form of the other arthropods discussed here. Insects Insect mouthparts exhibit a range of forms. The earliest insects had chewing mouthparts. Specialisation includes mouthparts modified for siphoning, piercing, sucking and sponging. These modifications have evolved a number of times. For example, mosquitoes (which are flies) and aphids (which are bugs) both ...
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