Muscomorpha
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Muscomorpha
The Brachyceran infraorder Muscomorpha is a large and diverse group of flies, containing the bulk of the Brachycera, and, most of the known flies. It includes a number of the most familiar flies, such as the housefly, the fruit fly, and the blow fly. The antennae are short, usually three-segmented, with a dorsal arista. Their bodies are often highly setose, and the pattern of setae is often taxonomically important. The larvae of muscomorphs (in the sense the name is used here; see below) have reduced head capsules, and the pupae are formed inside the exoskeleton of the last larval instar; exit from this puparium is by a circular line of weakness, and this pupal type is called "cyclorrhaphous"; this feature gives this group of flies their traditional name, Cyclorrhapha. Classification The name Cyclorrhapha is used, in various modern classifications, to represent either a subgroup within the infraorder Muscomorpha, or simply a rankless group within the Brachycera. In either ca ...
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Cyclorrhapha
Cyclorrhapha is an unranked taxon within the infraorder Muscomorpha. They are called "Cyclorrhapha" ('circular-seamed flies') with reference to the circular aperture through which the adult escapes the puparium. This is a circumscriptional name that has significant historical familiarity, but in the present classification, this name is synonymous with the more recent "Muscomorpha The Brachyceran infraorder Muscomorpha is a large and diverse group of flies, containing the bulk of the Brachycera, and, most of the known flies. It includes a number of the most familiar flies, such as the housefly, the fruit fly, and the blo ...". Cyclorrhapha underwent major adaptive radiation that led to the creation of over 72 000 species. These species share multiple attributes such as the 360-degree rotation of the male terminalia. References Muscomorpha Protostome unranked clades {{Muscomorpha-stub ...
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Asilomorpha
The Brachyceran infraorder Asilomorpha is a large and diverse group of flies, containing the bulk of the non muscoid Brachycera. The larvae of asilomorphs are extremely diverse in habits, as well. Classification In most modern classifications, the infraorder Asilomorpha is not recognized, as it is paraphyletic; the Empidoidea are the sister taxon to the Muscomorpha (used here in the same sense as the traditional Cyclorrhapha, not following the Tree of Life Web Project system in this one instance). However, these modern classifications are not Linnaean, and - while relatively easy to visualize on a phylogenetic tree diagram - do not lend themselves to any classification that uses Linnaean ranks (''see'' discussion ''for details''). Also, some classifications place the Nemestrinoidea Nemestrinoidea is a small, monophyletic superfamily of flies, whose relationship to the other Brachycera is uncertain; they are sometimes grouped with the Tabanomorpha rather than the Asilomorpha. Th ...
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Housefly
The housefly (''Musca domestica'') is a fly of the suborder Cyclorrhapha. It is believed to have evolved in the Cenozoic Era, possibly in the Middle East, and has spread all over the world as a commensal of humans. It is the most common fly species found in houses. Adults are gray to black, with four dark, longitudinal lines on the thorax, slightly hairy bodies, and a single pair of membranous wings. They have red eyes, set farther apart in the slightly larger female. The female housefly usually mates only once and stores the sperm for later use. She lays batches of about 100 eggs on decaying organic matter such as food waste, carrion, or feces. These soon hatch into legless white larvae, known as maggots. After two to five days of development, these metamorphose into reddish-brown pupae, about long. Adult flies normally live for two to four weeks, but can hibernate during the winter. The adults feed on a variety of liquid or semi-liquid substances, as well as solid ...
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Ephydroidea
The Ephydroidea are a superfamily of muscomorph flies, with over 6,000 species. Description A characteristic of adult Ephydroidea (shared with their relatives such as Calyptratae) is that the pedicel of the antenna has a dorsoventral seam or incision. Ecology Ephydroidea live in many habitats and have diverse diets. For example, most Ephydridae have larvae that are aquatic/semi-aquatic and feed as browsers or filter-feeders, but there are also species with terrestrial larvae that are egg predators, egg parasitoids, leaf miners or saprophages. Most Drosophilidae breed in rotting material where they feed on yeast and bacteria, but there are also species that attack whole fruits. Phylogeny A 2021 analysis found Ephydroidea to be the sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram ...
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Pupa
A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages thereof being egg, larva, pupa, and imago. The processes of entering and completing the pupal stage are controlled by the insect's hormones, especially juvenile hormone, prothoracicotropic hormone, and ecdysone. The act of becoming a pupa is called pupation, and the act of emerging from the pupal case is called eclosion or emergence. The pupae of different groups of insects have different names such as ''chrysalis'' for the pupae of butterflies and ''tumbler'' for those of the mosquito family. Pupae may further be enclosed in other structures such as cocoons, nests, or shells. Position in life cycle The pupal stage follows the larval stage and precedes adulthood (''imago'') in insects with complete metamorphosi ...
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Platypezoidea
The Platypezoidea are a superfamily of true flies of the section Aschiza. Their closest living relatives are the Syrphoidea, which, for example, contain the hoverflies. Like these, the adults do not burst open their pupal cases with a ptilinum when hatching, thus the Aschiza do not have the inverted-U-shaped suture above the antennae. They are, however, muscomorphs, thus have a particular type of pupal case resembling a rounded barrel and called puparium. Families Five families are placed in the Platypezoidea, listed below in taxonomic sequence: * Phoridae - coffin and scuttle flies (includes Sciadoceridae) * Opetiidae - flat-footed flies * Ironomyiidae - ironic flies * Lonchopteridae - spear-winged flies, pointed-wing flies * Platypezidae Platypezidae is a family of true flies of the superfamily Platypezoidea. The more than 250 species are found worldwide primarily in woodland habitats. A common name is flat-footed flies, but this is also used for the closely r ...
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Puparium
A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages thereof being egg, larva, pupa, and imago. The processes of entering and completing the pupal stage are controlled by the insect's hormones, especially juvenile hormone, prothoracicotropic hormone, and ecdysone. The act of becoming a pupa is called pupation, and the act of emerging from the pupal case is called eclosion or emergence. The pupae of different groups of insects have different names such as ''chrysalis'' for the pupae of butterflies and ''tumbler'' for those of the mosquito family. Pupae may further be enclosed in other structures such as cocoons, nests, or shells. Position in life cycle The pupal stage follows the larval stage and precedes adulthood (''imago'') in insects with complete metamorphosis ...
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Empidoidea
The Empidoidea are a large monophyletic superfamily of true flies, the sister taxon to the Muscomorpha (Cyclorrhapha). These two groups are sometimes united in the unranked taxon Eremoneura. There are some 10,000 known species within Empidoidea,Moulton & Wiegmann (2007) which are represented on all continents except Antarctica. They are known to have existed since the Jurassic period.Grimaldi & Engel (2005) Empidoidea has been subject to much debate regarding its phylogeny. Based on morphology alone, three major hypotheses had been proposed until 2007 and seemed to be consensus for some time—however, in 2018, Wahlberg & Johanson published the most current phylogeny supported by extensive genetic data, changing the relationship between quite a few families and tribes.Wahlberg & Johanson (2018) Description The majority of these insects are predatory, often with large compound eyes (sometimes covering almost the entire surface of the head), and tend to be associated with moist, t ...
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Aschiza
The Aschiza are a section (biology), section of the Brachycera. Two large families, the Syrphidae and the Phoridae, and a number of smaller taxa are in this group. They are similar to most of the familiar Muscomorpha with one notable exception; they do not possess a ptilinum, so lack the prominent ptilinal suture on the face as in other Schizophora, muscoid flies. They do still have a puparium with a circular emergence opening, but it is not as precisely ellipsoid in shape as is typical for other muscoids. The term was first used by Eduard Becher . References External links Tree of Life. Cyclorrhapha
Aschiza, {{Aschiza-stub ...
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Brachycera
The Brachycera are a suborder of the order Diptera. It is a major suborder consisting of around 120 families. Their most distinguishing characteristic is reduced antenna segmentation. Description A summary of the main physical characteristics is: * Antenna size (with eight or fewer flagellomeres) is reduced. * The maxillary palp (an elongated appendage near the mouth) has two segments or fewer. * The back portions of the larval head capsule extend into the prothorax (the anterior part of the thorax, which bears the first pair of legs). * Two distinct parts make up of the larval mandible (lower jaw). * The epandrium and hypandrium of the genitalia are separated in males. * No premandible is present on the lower surface of the labrum (the roof of the mouth). * The configuration of the CuA2 and A1 wing veins is distinct. Brachyceran flies can also be distinguished through behavior. Many of the species are predators or scavengers. Classification The structure of subgroups wit ...
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Syrphoidea
The Syrphoidea are a superfamily of flies containing only two families under present classification, one of which ( Syrphidae) has a great number of the most common and familiar flies. One of these familiar flies is ''Eristalis tenax ''Eristalis tenax'', the common drone fly, is a common, migratory, cosmopolitan species of hover fly. It is the most widely distributed syrphid species in the world, and is known from all regions except the Antarctic. It has been introduced in ...,'' or otherwise known as the drone fly. References External links Diptera superfamilies {{Syrphoidea-stub ...
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Opomyzoidea
The Opomyzoidea are a superfamily of flies. Biology Opomyzoids show a range of lifestyles including mining plant leaves (many Agromyzidae), feeding in grass stems ( Anthomyzidae and Opomyzidae), forming plant galls (''Fergusonina''), feeding on fungi (some Anthomyzidae and Asteiidae), feeding on sap flows of trees (some Aulacigastridae, Odiniidae and Periscelididae), living in galleries of wood-boring insects (Odiniidae) or in water-filled cavities of plants (phytotelmata; Aulacigastridae, Neurochaetidae and Periscelididae). However, the biology of most opomyzoid families is poorly known. Phylogeny The phylogeny of Opomyzoidea is controversial, with different authors assigning different families and different relationships among families. One study using molecular analysis concluded that the superfamily is not monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancesto ...
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