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Mesonychia
Mesonychia ("middle claws") is an extinct taxon of small- to large-sized carnivorous ungulates related to artiodactyls. Mesonychids first appeared in the early Paleocene, went into a sharp decline at the end of the Eocene, and died out entirely when the last genus, ''Mongolestes'', became extinct in the early Oligocene. In Asia, the record of their history suggests they grew gradually larger and more predatory over time, then shifted to scavenging and bone-crushing lifestyles before the group became extinct. Mesonychids probably originated in China, where the most primitive mesonychid, '' Yangtanglestes'', is known from the early Paleocene. They were also most diverse in Asia, where they occur in all major Paleocene faunas. Since other predators, such as creodonts and Carnivora, were either rare or absent in these animal communities, mesonychids most likely dominated the large predator niche in the Paleocene of eastern Asia. One genus, '' Dissacus'', had successfully spread to ...
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Mesonychidae
Mesonychidae (meaning "middle claws") is an extinct family of small to large-sized omnivorous-carnivorous mammals. They were endemic to North America and Eurasia during the Early Paleocene to the Early Oligocene, and were the earliest group of large carnivorous mammals in Asia. They are not closely related to any living mammals. Mesonychid taxonomy has long been disputed and they have captured popular imagination as "wolves on hooves," animals that combine features of both ungulates and carnivores. Skulls and teeth have similar features to early whales, and the family was long thought to be the ancestors of cetaceans. Recent fossil discoveries have overturned this idea; the consensus is that whales are highly derived artiodactyls. Some researchers now consider the family a sister group either to whales or to artiodactyls, close relatives rather than direct ancestors. Other studies define Mesonychia as basal to all ungulates, occupying a position between Perissodactyla and Ferae ...
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Paleocene
The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''palaiós'' meaning "old" and the Eocene Epoch (which succeeds the Paleocene), translating to "the old part of the Eocene". The epoch is bracketed by two major events in Earth's history. The K–Pg extinction event, brought on by an asteroid impact and possibly volcanism, marked the beginning of the Paleocene and killed off 75% of living species, most famously the non-avian dinosaurs. The end of the epoch was marked by the Paleocene–Eocene Thermal Maximum (PETM), which was a major climatic event wherein about 2,500–4,500 gigatons of carbon were released into the atmosphere and ocean systems, causing a spike in global temperatures and ocean acidification. In the Paleocene, the continents of the Northern Hemisphere were still connected ...
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Hapalodectidae
Hapalodectidae (literal translation 'soft biters': ('soft, tender'), ('biter')) is an extinct family of relatively small-bodied () mesonychian placental mammals from the Paleocene and Eocene of North America and Asia. Hapalodectids differ from the larger and better-known mesonychids by having teeth specialized for cutting (presumably meat), while the teeth of other mesonychids, such as ''Mesonyx'' or '' Sinonyx'', are more specialized for crushing bones. Hapalodectids were once considered a subfamily of the Mesonychidae, but the discovery of a skull of '' Hapalodectes hetangensis'' showed additional differences justifying placement in a distinct family. In particular, ''H. hetangensis'' has a postorbital bar closing the back of the orbit In celestial mechanics, an orbit is the curved trajectory of an object such as the trajectory of a planet around a star, or of a natural satellite around a planet, or of an artificial satellite around an object or position in space s ...
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Ungulate
Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. The term means, roughly, "being hoofed" or "hoofed animal". As a descriptive term, "ungulate" normally excludes cetaceans as they do not possess most of the typical morphological characteristics of other ungulates, but recent discoveries indicate that they were also descended from early artiodactyls. Ungulates are typically herbivorous and many employ specialized gut bacteria to allow them to digest cellulose. Some modern species, such as pigs, are omnivor ...
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Mongolestes
''Mongolestes'' ("Mongolian robber") is an extinct genus of mesonychid known from the 'Ulan Gochu' formation of Inner Mongolia, and likely originated in Asia. It was the last surviving representative of Mesonychia and became extinct in the early Oligocene. ''Mongolestes'' is distinct from other mesonychids in several dental features, including very large teeth and the loss of the third molar, and a mandibular symphysis In human anatomy, the facial skeleton of the skull the external surface of the mandible is marked in the median line by a faint ridge, indicating the mandibular symphysis (Latin: ''symphysis menti'') or line of junction where the two lateral halve ... that is steeper. Notes References * * Mesonychids Rupelian extinctions Eocene mammals of Asia Oligocene mammals of Asia Prehistoric placental genera {{paleo-mammal-stub ...
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Ankalagon
''Ankalagon saurognathus'' is an extinct carnivorous mammal of the family Mesonychidae, endemic to North America during the Paleocene epoch (63.3—60.2 mya), existing for approximately . Known from the Paleocene Nacimiento Formation of New Mexico, ''Ankalagon'' is the largest mesonychid known from the Paleocene of North America, and it provides the best evidence for sexual dimorphism in mesonychids. Description The main feature that distinguishes ''A. saurognathus'' from the ancestral '' Dissacus'' species is its size: ''Ankalagon'' grew to be as large as a bear,Paleocene mammals of the world
"Carnivores, creodonts and carnivorous ungulates: Mammals become predators"
as compared to the coyote or jackal-sized species of ''Dissacus''. In fact, the only North American mesonychids that surpassed '' ...
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Pachyaena
''Pachyaena'' (literally, "thick hyena") was a genus of heavily built, relatively short-legged mesonychids, early Cenozoic mammals that evolved before the origin of either modern hoofed animals or carnivores, and combined characteristics similar to both. The genus likely originated from Asia and spread to Europe, and from there to North America across a land bridge in what is now the North Atlantic ocean. Various described species of ''Pachyaena'' ranged in size from a coyote to a bear. However, recent work indicates that ''Pachyaena'' is paraphyletic, with ''P. ossifraga'' being closer to '' Synoplotherium'', '' Harpagolestes'' and ''Mesonyx'' than to ''P. gigantea''. Unlike many mesonychids, ''Pachyaena'' is known from skeletal material in addition to skulls and jaws. Analysis of three wolf- to bear-sized species from the early Eocene of Wyoming (Willwood formation) indicates they all had many adaptations for running, including paraxonic compressed feet with a vestigial fi ...
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Yangtanglestes
''Yantanglestes'' is a genus of small, Chinese mesonychid with slender jaws that first appeared during the Early Paleocene in the Thanetian stage. It was found throughout Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are .... It is the oldest known mesonychid. ''Yantanglestes'' became extinct during the Nongshanian division of the Late Paleocene,Missiaen, Pieter. "亚洲早古近纪哺乳动物生物年代学与生物地理学的新认识."古脊椎动物学报 49.1 (2011)paper/ref> and lived sympatrically with its descendant genera, including '' Dissacus'', '' Sinonyx'' and '' Jiangxia''. References External linksScience Mesonychids Paleocene mammals Thanetian life Paleocene mammals of Asia Paleogene China Prehistoric animals of China Fossil taxa described in 1980 ...
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Dissacus
''Dissacus'' is a genus of extinct carnivorous jackal to coyote-sized mammals within the family Mesonychidae, an early group of hoofed mammals that evolved into hunters and omnivores. Their fossils are found in Paleocene to Early Eocene aged strata in France, Asia and southwest North America, from 66 to 50.3 mya, existing for approximately . Orientation patch analysis of the molar teeth of the North American ''D. praenuntius'' suggests it was an omnivore that ate a lot of meat, not an exclusive meat-eater like a cat or weasel. It shared its environment with more omnivorous mammals of a similar body size. Though they are not ancestral to Carnivora, ''Dissacus'' species may have had similar roles in Paleocene-early Eocene environments as the foxes and other small canids that evolved later: generalized hunters who also ate fruit or other foods, and caught small animals that lived on the ground. The bear-sized ''Ankalagon'' is closely related to ''Dissacus''. Some paleontologi ...
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Artiodactyl
The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, odd-toed ungulates bear weight on an odd number of the five toes. Another difference between the two is that many other even-toed ungulates (with the exception of Suina) digest plant cellulose in one or more stomach chambers rather than in their intestine as the odd-toed ungulates do. Cetaceans (whales, dolphins, and porpoises) evolved from even-toed ungulates, and are therefore often classified under the same taxonomic branch because a species cannot outgrow its evolutionary ancestry; some modern taxonomists combine the two under the name Cetartiodactyla , while others opt to include cetaceans in the already-existing Artiodactyla. The roughly 270 land-based even-toed ungulate species include pigs, peccaries, hippopo ...
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Triisodontidae
Triisodontidae is an extinct, probably paraphyletic, or possibly invalid family of mesonychian placental mammals. Most triisodontid genera lived during the Paleocene in North America, but the genus '' Andrewsarchus'' (if it is a mesonychian, and not an artiodactyl) is known from the middle Eocene of Asia. Triisodontids were the first relatively large predatory mammals to appear in North America following the extinction of the non-bird dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...s. They differ from other mesonychian families in having less highly modified teeth. Because of their comparatively simpler teeth, the Triisodontids are regarded as basal mesonychids. A recent study found them to be a paraphyletic assemblage of stem-mesonychians.Sarah L. Shelley, Thomas E ...
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Creodonts
Creodonta ("meat teeth") is a former order of extinct carnivorous placental mammals that lived from the early Paleocene to the late Miocene epochs in North America, Europe, Asia and Africa. Originally thought to be a single group of animals ancestral to the modern Carnivora, this order is now usually considered a polyphyletic assemblage of two different groups, the Oxyaenids and the Hyenodonts, not a natural group. Oxyaenids are first known from the Palaeocene of North America, while hyaenodonts hail from the Palaeocene of Africa. Creodonts were the dominant carnivorous mammals from , peaking in diversity and prevalence during the Eocene. The first large, obviously carnivorous mammals appeared with the radiation of the oxyaenids in the late Paleocene. During the Paleogene, "creodont" species were the most abundant terrestrial carnivores in the Old World. In Oligocene Africa, hyaenodonts were the dominant group of large flesh-eaters, persisting until the middle of the Miocene. ...
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