Metastoma
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Metastoma
The metastoma is a ventral single plate located in the opisthosoma of non-arachnid dekatriatan chelicerates such as eurypterids, chasmataspidids and the genus ''Houia''. The metastoma located between the base of 6th prosomal appendage pair and may had functioned as part of the animal's feeding structures. It most likely represented a fused appendage pair originated from somite 7 (first opisthosomal segment), thus homologous to the chilaria of horseshoe crab and 4th walking leg pair of sea spider. In eurypterids, the plate was typically cordate (heart-shaped) in shape, though differed in shape in some genera, such as ''Megalograptus ''Megalograptus'' is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of ''Megalograptus'' have been recovered in deposits of Katian (Late Ordovician) age in North America. The genus contains five species: ''M. alveolatu ...''. References Chelicerate anatomy Dekatriata {{Chelicerata-stub ...
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Eurypterid
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 Myr, million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic Chelicerata, chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Tri ...
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Eurypterus Anatomy
''Eurypterus'' ( ) is an extinct genus of eurypterid, a group of organisms commonly called "sea scorpions". The genus lived during the Silurian period, from around 432 to 418 million years ago. ''Eurypterus'' is by far the most well-studied and well-known eurypterid. ''Eurypterus'' fossil specimens probably represent more than 95% of all known eurypterid specimens. There are fifteen species belonging to the genus ''Eurypterus'', the most common of which is ''E. remipes'', the first eurypterid fossil discovered and the state fossil of New York. Members of ''Eurypterus'' averaged at about in length, but the largest individual discovered was estimated to be long. They all possessed spine-bearing appendages and a large paddle they used for swimming. They were generalist species, equally likely to engage in predation or scavenging. Discovery The first fossil of ''Eurypterus'' was found in 1818 by S. L. Mitchill, a fossil collector. It was recovered from the Bertie Formation of ...
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Chasmataspidid
Chasmataspidids, sometime referred to as chasmataspids, are a group of extinct chelicerate arthropods that form the order Chasmataspidida. Chasmataspidids are probably related to horseshoe crabs (Xiphosura) and/or sea scorpions (Eurypterida), with more recent studies suggest that they form a clade (Dekatriata) with Eurypterida and Arachnida. Chasmataspidids are known sporadically in the fossil record through to the mid-Devonian, with possible evidence suggesting that they were also present during the late Cambrian. Chasmataspidids are most easily recognised by having an opisthosoma divided into a wide forepart (preabdomen) and a narrow hindpart (postabdomen) each comprising 4 and 9 segments respectively. There is some debate about whether they form a natural (i.e. monophyletic) group. Distribution Chasmataspidids survived at least since Ordovician to mid-Devonian in age. As of 2019, most chasmataspidids (with a total of 9 species) are known from the Devonian strata, while the p ...
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Houia
''Houia'' is an extinct genus of dekatriatan, a clade of chelicerate arthropods. Fossils of ''Houia'' have been discovered in deposits of the Early Devonian period in Guangxi and Yunnan, both in China. The genus contains two species: ''H. guangxiensis'', from the Pragian to Emsian epoch of Guangxi; and ''H. yueya'', the type species, from the Lochkovian epoch of Yunnan. The name of the genus is derived from the Chinese character 鲎 (''hòu''), meaning "horseshoe crab". ''Houia yueya'' was originally described as a species of the xiphosuran (horseshoe crab) genus '' Kasibelinurus'' (''Kasibelinurus yueya'') in 2013, with its narrow opisthosoma (the trunk section) being misinterpreted as incompletely preserved (lacking lateral regions). The species name ''yueya'' comes from the Chinese characters 月 (''yuè'', meaning "Moon") and 牙 (''yá'', meaning "crescent"), referring to the crescentic shape of its carapace (the dorsal plate of the prosoma or head). ''H. yueya'' ...
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Megalograptus
''Megalograptus'' is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of ''Megalograptus'' have been recovered in deposits of Katian (Late Ordovician) age in North America. The genus contains five species: ''M. alveolatus'', ''M. ohioensis'', ''M. shideleri'', ''M. welchi'' and ''M. williamsae'', all based on fossil material found in the United States. Fossils unassigned to any particular species have also been found in Canada. The generic name translates to "great writing" and originates from the mistaken original belief that ''Megalograptus'' was a type of graptolite, often given names ending with -''graptus'' (meaning 'writing'). ''Megalograptus'' was a large predatory megalograptid eurypterid, with the largest and best known species, ''M. ohioensis'', reaching body lengths of . Some species were substantially smaller, with the smallest, belonging to a hitherto undetermined species, only growing to about in length. Morphologically, ''Megalograptus'' ...
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Ventral
Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. This position provides a definition of what is at the front ("anterior"), behind ("posterior") and so on. As part of defining and describing terms, the body is described through the use of anatomical planes and anatomical axis, anatomical axes. The meaning of terms that are used can change depending on whether an organism is bipedal or quadrupedal. Additionally, for some animals such as invertebrates, some terms may not have any meaning at all; for example, an animal that is radially symmetrical will have no anterior surface, but can still have a description that a part is close to the middle ("proximal") or further from the middle ("distal"). International organisations have determined vocabularies that are often used as standard vocabular ...
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Opisthosoma
The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma (cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects to an abdomen (and is often referred to as such), the opisthosoma is differentiated by its inclusion of the respiratory organs (book lungs or book gills) and the heart. Segments The number of segments and appendages on the opisthosoma vary. Scorpions have 13, but the first is only seen during its embryological development. Other arachnids have fewer; harvestmen, for instance, have only ten. In general, appendages are absent or reduced, although in horseshoe crabs they persist as large plate-like limbs, called opercula or branchiophores, bearing the book gills, and that function in locomotion and gas exchange. In most chelicerates the opisthosomal limbs are greatly reduced and persist only as specialized structures, such as the silk-producing ...
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Arachnid
Arachnida () is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons. Almost all adult arachnids have eight legs, although the front pair of legs in some species has converted to a sensory function, while in other species, different appendages can grow large enough to take on the appearance of extra pairs of legs. The term is derived from the Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, who was turned into a spider. Almost all extant arachnids are terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 100,000 named species, of which 47,000 are species of spiders. Morphology Almost all adult arachnids have eight legs, unlike adult inse ...
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Dekatriata
Dekatriata is a clade of planatergan chelicerates including the groups Arachnida, Chasmataspidida, Eurypterid Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is l ...a and additionally two stem-genera '' Winneshiekia'' and '' Houia''. Dekatriata is defined by an opisthosoma with 13 segments as groundplan (the number proposed to be secondarily reduced in most arachnid orders) and fused, plate-like appendages on the first opisthosomal segment (somite VII). References Middle Ordovician first appearances {{Chelicerata-stub ...
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Chelicerates
The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. The Chelicerata originated as marine animals in the Middle Cambrian period; the first confirmed chelicerate fossils, belonging to ''Sanctacaris'', date from 508 million years ago. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Zoologica Scripta
''Zoologica Scripta'' is a bimonthly peer-reviewed scientific journal on systematic zoology, published by Wiley-Blackwell on behalf of the Norwegian Academy of Science and Letters and the Royal Swedish Academy of Sciences. It was established in 1972. The current chief editor is Per Sundberg. According to the ''Journal Citation Reports'', the journal has a 2020 impact factor The impact factor (IF) or journal impact factor (JIF) of an academic journal is a scientometric index calculated by Clarivate that reflects the yearly mean number of citations of articles published in the last two years in a given journal, as i ... of 3.140, ranking it 12th out of 174 journals in the category "Zoology". See also * '' Arkiv för Zoologi'' References External links * * Zoology journals Royal Swedish Academy of Sciences Publications established in 1972 Bimonthly journals Wiley-Blackwell academic journals Academic journals associated with learned and professional societies
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