Maniraptoran
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Maniraptoran
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Deinonychosauria, Oviraptorosauria and Therizinosauria. ''Ornitholestes'' and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period (see ''Eshanosaurus''), and survive today as living birds. Description Maniraptorans are characterized by long arms and three-fingered hands (though reduced or fused in some lineages), as well as a "half-moon shaped" (semi-lunate) bone in the wrist ( carpus). In 2004, Tom Holtz and Halszka Osmólska pointed out six other maniraptoran characters relating to specific details of the skeleton. Unlike most other saurischian dinosaurs, whi ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Therizinosauria
Therizinosaurs (once called segnosaurs) were large herbivorous theropod dinosaurs whose fossils have been found across the Early to Late Cretaceous deposits in Asia and North America. Various features of the forelimbs, skull and pelvis unite these finds as both theropods and maniraptorans, close relatives to birds. The name of the representative genus, ''Therizinosaurus'', is derived from the Greek (, 'to reap' or 'scythe')Translated paper
and (, 'lizard'). The older representative, ''Segnosaurus'', is derived from the ('slow') and the Greek .


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Jurassic Period
The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of the Mesozoic Era and is named after the Jura Mountains, where limestone strata from the period were first identified. The start of the Jurassic was marked by the major Triassic–Jurassic extinction event, associated with the eruption of the Central Atlantic Magmatic Province. The beginning of the Toarcian Stage started around 183 million years ago and is marked by an extinction event associated with widespread oceanic anoxia, ocean acidification, and elevated temperatures likely caused by the eruption of the Karoo-Ferrar large igneous provinces. The end of the Jurassic, however, has no clear boundary with the Cretaceous and is the only boundary between geological periods to remain formally undefined. By the beginning of the Jurassic, t ...
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Maniraptoriformes
Maniraptoriformes is a clade of dinosaurs with pennaceous feathers and wings that contains ornithomimosaurs and maniraptorans. This group was named by Thomas Holtz, who defined it as "the most recent common ancestor of '' Ornithomimus'' and birds, and all descendants of that common ancestor." Fossil evidence Many fossils have been discovered in recent years, particularly in China. Many of the feathered dinosaurs belong to this clade. In particular, a fossil of the Alvarezsauridae '' Shuvuuia'' has a version of keratin consistent with that of avian feathers. Many nearly complete fossil maniraptoriforms are known from the late Jurassic. ''Archaeopteryx'' is known from Bavaria at 155–150 Ma. '' Ornitholestes'', the troodontid ''Hesperornithoides'', '' Coelurus fragilis'' and '' Tanycolagreus topwilsoni'' are all known from the Morrison Formation in Wyoming at about 150 Ma. The Daohugou biota, including ''Anchiornis'' and ''Epidexipteryx'', is the earliest record of man ...
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Ornithomimosauria
Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period (geology), Period of Laurasia (now Asia, Europe and North America), as well as Africa and possibly Australia. The group first appeared in the Early Cretaceous and persisted until the Late Cretaceous. Primitive members of the group include ''Nqwebasaurus'', ''Pelecanimimus'', ''Shenzhousaurus'', ''Hexing'' and ''Deinocheirus'', the arms of which reached 2.4 m (8 feet) in length. More advanced species, members of the family Ornithomimidae, include ''Gallimimus'', ''Struthiomimus'', and ''Ornithomimus''. Some paleontologists, like Paul Sereno, consider the enigmatic Alvarezsauridae, alvarezsaurids to be close relatives of the ornithomimosaurs and place them together in the superfamily Ornithomimoidea (see classification below). Description The skulls of ornithomimosaurs ...
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Ornitholestes
''Ornitholestes'' (meaning "bird robber") is a small theropod dinosaur of the late Jurassic (Brushy Basin Member of the Morrison Formation, middle Kimmeridgian age, about 154 million years agoTurner, C.E. and Peterson, F., (1999). "Biostratigraphy of dinosaurs in the Upper Jurassic Morrison Formation of the Western Interior, U.S.A." Pp. 77–114 in Gillette, D.D. (ed.), ''Vertebrate Paleontology in Utah''. Utah Geological Survey Miscellaneous Publication 99-1.) of Western Laurasia (the area that was to become North America). To date, ''Ornitholestes'' is known only from a single partial skeleton with a badly crushed skull found at the Bone Cabin Quarry near Medicine Bow, Wyoming, in 1900. It was described by Henry Fairfield Osborn in 1903. An incomplete hand was later attributed to ''Ornitholestes'', although it now appears to belong to ''Tanycolagreus''. The type (and only known) species is ''O. hermanni''. The specific name honors the American Museum of Natural History prepar ...
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Oviraptorosauria
Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from ''Caudipteryx'', which was the size of a turkey, to the 8-meter-long, 1.4-ton ''Gigantoraptor''. The group (along with all maniraptoran dinosaurs) is close to the ancestry of birds. Some researchers such as Maryanska ''et al'' (2002) and Osmólska ''et al.'' (2004) have proposed that they may represent primitive flightless birds.Osmólska, Halszka, Currie, Philip J., Brasbold, Rinchen (2004) "The Dinosauria" Weishampel, Dodson, Osmólska. "Chapter 8 Oviraptorosauria" University of California Press. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is sparse.Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) ...
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Deinonychosauria
Deinonychosauria is a clade of paravian dinosaurs which lived from the Late Jurassic to the Late Cretaceous periods. Fossils have been found across the globe in North America, Europe, Africa, Asia, South America, and Antarctica,Case, J.A., Martin, J.E., and Reguero, M. (2007). "A dromaeosaur from the Maastrichtian of James Ross Island and the Late Cretaceous Antarctic dinosaur fauna." Pp. 1–4 in Cooper, A., Raymond, C., and Team, I.E. (eds.), ''Antarctica: a Keystone in a Changing World – Online Proceedings for the Tenth International Symposium on Antarctic Earth Sciences'', U.S. Geological Survey Open-File Report 2007-1047, SRP 083. U.S. Geological Survey, Washington, D.C. with fossilized teeth giving credence to the possibility that they inhabited Australia as well. This group of dinosaurs are known for their sickle-shaped toe claws and features in the shoulder bones. Deinonychosauria is commonly defined as all dinosaurs more closely related to dromaeosaurids (such as ''Dein ...
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Avialae
Avialae ("bird wings") is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used (see below). ''Archaeopteryx lithographica'', from the late Jurassic Period Solnhofen Formation of Germany, is possibly the earliest known avialan which may have had the capability of powered flight, though it might have been a deinonychosaur instead. Several older (but non flight-capable) avialans are known from the late Jurassic Tiaojishan Formation of China, dated to about 160 million years ago. Definition Most researchers define Avialae as branch-based clade, though definitions vary. Many authors have used a definition similar to "all theropods closer to birds than to ''Deinonychus''."Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.) (2004). ''The Dinosauria'', Second Edition. University of California Press., 861 pp. ...
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Ornithomimus Velox
''Ornithomimus'' (; "bird mimic") is a genus of ornithomimid dinosaurs from the Late Cretaceous Period of what is now North America. ''Ornithomimus'' was a swift bipedal theropod which fossil evidence indicates was covered in feathers, equipped with a small toothless beak that may indicate an omnivorous diet. It is usually classified into two species: the type species, ''Ornithomimus velox'', and a referred species, ''Ornithomimus edmontonicus''. ''O. velox'' was named in 1890 by Othniel Charles Marsh on the basis of a foot and partial hand from the late Maastrichtian-age Denver Formation of Colorado, United States. Another seventeen species have been named since, though most of them have subsequently been assigned to new genera or shown to be not directly related to ''Ornithomimus velox''. The best material of species still considered part of the genus has been found in Alberta, Canada, representing the species ''O. edmontonicus'', known from several skeletons from the early Maa ...
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Bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the bee hummingbird to the ostrich. There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming. B ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, are among ...
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