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Macroevolutionary
Macroevolution usually means the evolution of large-scale structures and traits that go significantly beyond the intraspecific variation found in microevolution (including speciation). In other words, macroevolution is the evolution of taxa above the species level (genera, families, orders, etc.). Macroevolution is often thought to require the evolution of completely new structures such as entirely new organs. However, fundamentally novel structures are not necessary for dramatic evolutionary change. For instance, the evolution of mammal diversity in the past 100 million years has not required any major innovation. All of this diversity can be explained by modification of existing organs. In most cases, macroevolution cannot be observed directly because numerous mutations are required for large-scale changes. Hence specific methods are required to study and prove it. However, in rare cases single mutations can cause dramatic change and thus serve as models for macro-evolutionar ...
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Evolution
Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation tends to exist within any given population as a result of genetic mutation and recombination. Evolution occurs when evolutionary processes such as natural selection (including sexual selection) and genetic drift act on this variation, resulting in certain characteristics becoming more common or more rare within a population. The evolutionary pressures that determine whether a characteristic is common or rare within a population constantly change, resulting in a change in heritable characteristics arising over successive generations. It is this process of evolution that has given rise to biodiversity at every level of biological organisation, including the levels of species, individual organisms, and molecules. The theory of evolution by ...
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Richard Goldschmidt
Richard Benedict Goldschmidt (April 12, 1878 – April 24, 1958) was a German-born American geneticist. He is considered the first to attempt to integrate genetics, development, and evolution. He pioneered understanding of reaction norms, genetic assimilation, dynamical genetics, sex determination, and heterochrony.Dietrich, Michael R. (2003)Richard Goldschmidt: hopeful monsters and other 'heresies.'''Nature Reviews Genetics'' 4 (Jan.): 68-74. Controversially, Goldschmidt advanced a model of macroevolution through macromutations popularly known as the "Hopeful Monster" hypothesis. Goldschmidt also described the nervous system of the nematode, a piece of work that influenced Sydney Brenner to study the wiring diagram of ''Caenorhabditis elegans'', winning Brenner and his colleagues the Nobel Prize in 2002. Childhood and education Goldschmidt was born in Frankfurt-am-Main, Germany to upper-middle class parents of Ashkenazi Jewish heritage. He had a classical education and ...
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Protein
Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, responding to stimuli, providing structure to cells and organisms, and transporting molecules from one location to another. Proteins differ from one another primarily in their sequence of amino acids, which is dictated by the nucleotide sequence of their genes, and which usually results in protein folding into a specific 3D structure that determines its activity. A linear chain of amino acid residues is called a polypeptide. A protein contains at least one long polypeptide. Short polypeptides, containing less than 20–30 residues, are rarely considered to be proteins and are commonly called peptides. The individual amino acid residues are bonded together by peptide bonds and adjacent amino acid residues. The sequence of amino acid residue ...
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Bone
A bone is a Stiffness, rigid Organ (biology), organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red blood cell, red and white blood cells, store minerals, provide structure and support for the body, and enable animal locomotion, mobility. Bones come in a variety of shapes and sizes and have complex internal and external structures. They are lightweight yet strong and hard and serve multiple Function (biology), functions. Bone tissue (osseous tissue), which is also called bone in the mass noun, uncountable sense of that word, is hard tissue, a type of specialized connective tissue. It has a honeycomb-like matrix (biology), matrix internally, which helps to give the bone rigidity. Bone tissue is made up of different types of bone cells. Osteoblasts and osteocytes are involved in the formation and mineralization (biology), mineralization of bone; osteoclasts are involved in the bone resorption, resor ...
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Vein
Veins are blood vessels in humans and most other animals that carry blood towards the heart. Most veins carry deoxygenated blood from the tissues back to the heart; exceptions are the pulmonary and umbilical veins, both of which carry oxygenated blood to the heart. In contrast to veins, arteries carry blood away from the heart. Veins are less muscular than arteries and are often closer to the skin. There are valves (called ''pocket valves'') in most veins to prevent backflow. Structure Veins are present throughout the body as tubes that carry blood back to the heart. Veins are classified in a number of ways, including superficial vs. deep, pulmonary vs. systemic, and large vs. small. * Superficial veins are those closer to the surface of the body, and have no corresponding arteries. *Deep veins are deeper in the body and have corresponding arteries. *Perforator veins drain from the superficial to the deep veins. These are usually referred to in the lower limbs and feet. *Communic ...
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Heart
The heart is a muscular organ in most animals. This organ pumps blood through the blood vessels of the circulatory system. The pumped blood carries oxygen and nutrients to the body, while carrying metabolic waste such as carbon dioxide to the lungs. In humans, the heart is approximately the size of a closed fist and is located between the lungs, in the middle compartment of the chest. In humans, other mammals, and birds, the heart is divided into four chambers: upper left and right atria and lower left and right ventricles. Commonly the right atrium and ventricle are referred together as the right heart and their left counterparts as the left heart. Fish, in contrast, have two chambers, an atrium and a ventricle, while most reptiles have three chambers. In a healthy heart blood flows one way through the heart due to heart valves, which prevent backflow. The heart is enclosed in a protective sac, the pericardium, which also contains a small amount of fluid. The wall ...
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Fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of living fish species are ray-finned fish, belonging to the class Actinopterygii, with around 99% of those being teleosts. The earliest organisms that can be classified as fish were soft-bodied chordates that first appeared during the Cambrian period. Although they lacked a true spine, they possessed notochords which allowed them to be more agile than their invertebrate counterparts. Fish would continue to evolve through the Paleozoic era, diversifying into a wide variety of forms. Many fish of the Paleozoic developed external armor that protected them from predators. The first fish with jaws appeared in the Silurian period, after which many (such as sharks) became formidable marine predators rather than just the prey of arthropods. Mos ...
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Swim Bladder
The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled Organ (anatomy), organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth without having to expend energy in swimming. Also, the Dorsum (biology), dorsal position of the swim bladder means the center of mass is below the centroid, center of volume, allowing it to act as a stabilizing agent. Additionally, the swim bladder functions as a resonator, resonating chamber, to produce or receive sound. The swim bladder is evolutionarily Homology (biology), homologous to the lungs. Charles Darwin remarked upon this in ''On the Origin of Species''.Darwin, Charles (1859''Origin of Species''Page 190, reprinted 1872 by D. Appleton. Darwin reasoned that the lung in air-breathing vertebrates had derived from a more primitive swim bladder. In the embryonic stages, some species, such as Ophioblennius atlanticus, ...
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Lung
The lungs are the primary organs of the respiratory system in humans and most other animals, including some snails and a small number of fish. In mammals and most other vertebrates, two lungs are located near the backbone on either side of the heart. Their function in the respiratory system is to extract oxygen from the air and transfer it into the bloodstream, and to release carbon dioxide from the bloodstream into the atmosphere, in a process of gas exchange. Respiration is driven by different muscular systems in different species. Mammals, reptiles and birds use their different muscles to support and foster breathing. In earlier tetrapods, air was driven into the lungs by the pharyngeal muscles via buccal pumping, a mechanism still seen in amphibians. In humans, the main muscle of respiration that drives breathing is the diaphragm. The lungs also provide airflow that makes vocal sounds including human speech possible. Humans have two lungs, one on the left and on ...
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Reptile Scale
Reptile skin is covered with scutes or scales which, along with many other characteristics, distinguish reptiles from animals of other classes. They are made of alpha and beta-keratin and are formed from the epidermis (contrary to fish, in which the scales are formed from the dermis). The scales may be ossified or tubercular, as in the case of lizards, or modified elaborately, as in the case of snakes. Smith, Malcolm A. (1931). ''The Fauna of British India, Ceylon and Burma. Vol. I.—Loricata, Testudines.'' London: Secretary of State for India in Council. (Taylor and Francis, printers). xxviii + 185 pp. + Plates I-II. ("Skin", p. 30). The scales on the top of lizard and snake heads has also been called ''pileus'', after the Latin word for cap, referring to the fact that these scales sit on the skull like a cap. Lizard scales Lizard scales vary in form from tubercular to platelike, or imbricate (overlapping). These scales, which on the surface are composed of horny (keratinized) ...
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Feather
Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and a premier example of a complex evolutionary novelty. They are among the characteristics that distinguish the extant birds from other living groups. Although feathers cover most of the bird's body, they arise only from certain well-defined tracts on the skin. They aid in flight, thermal insulation, and waterproofing. In addition, coloration helps in communication and protection. Plumology (or plumage science) is the name for the science that is associated with the study of feathers. Feathers have a number of utilitarian, cultural, and religious uses. Feathers are both soft and excellent at trapping heat; thus, they are sometimes used in high-class bedding, especially pillows, blankets, and mattresses. They are also used as filling for winter cloth ...
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Bird Wing
Bird wings are a paired forelimb in birds. The wings give the birds the Bird flight, ability to fly, creating Lift (force), lift. Terrestrial flightless birds have reduced wings or none at all (for example, moa). In Waterfowl, aquatic flightless birds (penguins), wings can serve as Flipper (anatomy), flippers. Anatomy Like most other tetrapods, the forelimb of birds consists of the shoulder (with the humerus), the forearm (with the ulna and the Radius (bone), radius), and the hand. The hand of birds is substantially transformed: some of its bones have been reduced, and some others have merged with each other. Three bones of the metacarpus and part of the carpal bones merge into a carpometacarpus. The bones of three fingers are attached to it. The frontmost one bears an alula—a group of feathers that act like the slats of an airplane. Usually, this finger has one phalanx bone, the next has two, and the back has one (but some birds have one more phalanx on the first two fing ...
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