Lycopod
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Lycopod
Lycopodiopsida is a class of vascular plants known as lycopods, lycophytes or other terms including the component lyco-. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching stems bearing simple leaves called microphylls and reproduce by means of spores borne in sporangia on the sides of the stems at the bases of the leaves. Although living species are small, during the Carboniferous, extinct tree-like forms formed huge forests that dominated the landscape and contributed to coal deposits. The nomenclature and classification of plants with microphylls varies substantially among authors. A consensus classification for extant (living) species was produced in 2016 by the Pteridophyte Phylogeny Group (PPG I), which places them all in the class Lycopodiopsida, which includes the classes Isoetopsida and Selaginellopsida used in other systems. (See Table 2.) Alternative classification systems have used ranks fro ...
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Fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Lycopodiales
The Lycopodiaceae (class Lycopodiopsida, order Lycopodiales) are an old family of vascular plants, including all of the core clubmosses and firmosses, comprising 16 accepted genera and about 400 known species. This family originated about 380 million years ago in the early Devonian, though the diversity within the family has been much more recent. "Wolf foot" is another common name for this family due to the resemblance of either the roots or branch tips to a wolf's paw. Description Members of Lycopodiaceae are not spermatophytes and so do not produce seeds. Instead they produce spores, which are oily and flammable, and are the most economically important aspects of these plants. The spores are of one size (i.e. the plants are Spore#Classification_of_spore-producing_organisms, isosporous) and are borne on a specialized structure at the apex of a shoot called a strobilus (plural: strobili), which resembles a tiny Club (weapon), battle club, from which the common name derives. Membe ...
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Zosterophyll
The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosterophyllophyta or Zosterophyta and the class or plesion Zosterophyllopsida or Zosteropsida. They were among the first vascular plants in the fossil record, and had a world-wide distribution. They were probably stem-group lycophytes, forming a sister group to the ancestors of the living lycophytes. By the late Silurian (late Ludlovian, about ) a diverse assemblage of species existed, examples of which have been found fossilised in what is now Bathurst Island in Arctic Canada. Morphology The stems of zosterophylls were either smooth or covered with small spines known as enations, branched dichotomously, and grew at the ends by unrolling, a process known as circinate vernation. The stems had a central vascular column in which the protoxylem ...
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Euphyllophyte
The euphyllophytes are a clade of plants within the tracheophytes (the vascular plants). The group may be treated as an unranked clade, a division under the name Euphyllophyta or a subdivision under the name Euphyllophytina. The euphyllophytes are characterized by the possession of true leaves ("megaphylls"), and comprise one of two major lineages of extant vascular plants. As shown in the cladogram below, the euphyllophytes have a sister relationship to the lycopodiophytes or lycopsids. Unlike the lycopodiophytes, which consist of relatively few presently living or extant taxa, the euphyllophytes comprise the vast majority of vascular plant lineages that have evolved since both groups shared a common ancestor more than 400 million years ago. The euphyllophytes consist of two lineages, the spermatophytes A spermatophyte (; ), also known as phanerogam (taxon Phanerogamae) or phaenogam (taxon Phaenogamae), is any plant that produces seeds, hence the alternative name seed plant. Sp ...
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Selaginellales
''Selaginella'' is the sole genus of vascular plants in the family Selaginellaceae, the spikemosses or lesser clubmosses. This family is distinguished from Lycopodiaceae (the clubmosses) by having scale-leaves bearing a ligule and by having spores of two types. They are sometimes included in an informal paraphyletic group called the "fern allies". '' S. moellendorffii'' is an important model organism. Its genome has been sequenced by the United States Department of Energy's Joint Genome Institute. The name ''Selaginella'' was erected by Palisot de Beauvois solely for the species ''Selaginella selaginoides'', which turns out (with the closely related ''Selaginella deflexa'') to be a clade that is sister to all other ''Selaginellas'', so any definitive subdivision of the species into separate genera leaves two taxa in ''Selaginella'', with the hundreds of other species in new or resurrected genera. ''Selaginella'' occurs mostly in the tropical regions of the world, with a hand ...
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Selaginella
''Selaginella'' is the sole genus of vascular plants in the family Selaginellaceae, the spikemosses or lesser clubmosses. This family is distinguished from Lycopodiaceae (the clubmosses) by having scale-leaves bearing a ligule and by having spores of two types. They are sometimes included in an informal paraphyletic group called the "fern allies". '' S. moellendorffii'' is an important model organism. Its genome has been sequenced by the United States Department of Energy's Joint Genome Institute. The name ''Selaginella'' was erected by Palisot de Beauvois solely for the species ''Selaginella selaginoides'', which turns out (with the closely related ''Selaginella deflexa'') to be a clade that is sister to all other ''Selaginellas'', so any definitive subdivision of the species into separate genera leaves two taxa in ''Selaginella'', with the hundreds of other species in new or resurrected genera. ''Selaginella'' occurs mostly in the tropical regions of the world, with a hand ...
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Isoetes
''Isoetes'', commonly known as the quillworts, is the only extant genus of plants in the family Isoetaceae, which is in the class of lycopods. There are currently 192 recognized species, with a cosmopolitan distribution but with the individual species often scarce to rare. Some botanists split the genus, separating two South American species into the genus ''Stylites'', although molecular data place these species among other species of ''Isoetes'', so that ''Stylites'' does not warrant taxonomic recognition. Species of ''Isoetes'' virtually identical to modern forms have existed since the Jurassic epoch. The name of the genus may also be spelled ''Isoëtes''. The diaeresis (two dots over the e) indicates that the o and the e are to be pronounced in two distinct syllables. Including this in print is optional; either spelling (''Isoetes'' or ''Isoëtes'') is correct. Description Quillworts are mostly aquatic or semi-aquatic in clear ponds and slow-moving streams, though several ( ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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Gametophyte
A gametophyte () is one of the two alternation of generations, alternating multicellular organism, multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the Sexual reproduction of plants, sexual phase in the life cycle of plants and algae. It develops sex organs that produce gametes, haploid sex cells that participate in fertilization to form a diploid zygote which has a double set of chromosomes. Cell division of the zygote results in a new diploid multicellular organism, the second stage in the life cycle known as the sporophyte. The sporophyte can produce haploid spores by meiosis that on germination produce a new generation of gametophytes. Algae In some multicellular green algae (''Ulva lactuca'' is one example), red algae and brown algae, sporophytes and gametophytes may be externally indistinguishable (isomorphic). In ''Ulva (genus), Ulv ...
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Myco-heterotrophy
Myco-heterotrophy (from Ancient Greek, Greek μύκης , "fungus", ἕτερος ', "another", "different" and τροφή ', "nutrition") is a symbiosis, symbiotic relationship between certain kinds of plants and fungi, in which the plant gets all or part of its food from parasitism upon fungi rather than from photosynthesis. A myco-heterotroph is the parasitic plant partner in this relationship. Myco-heterotrophy is considered a kind of Cheating (biology), cheating relationship and myco-heterotrophs are sometimes informally referred to as "mycorrhizal cheaters". This relationship is sometimes referred to as mycotrophy, though this term is also used for plants that engage in Mutualism (biology), mutualistic mycorrhizal relationships. Relationship between myco-heterotrophs and host fungi Full (or obligate) myco-heterotrophy exists when a non-photosynthetic plant (a plant largely lacking in chlorophyll or otherwise lacking a functional photosystem) gets all of its food from the ...
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Sporophyte
A sporophyte () is the diploid multicellular stage in the life cycle of a plant or alga which produces asexual spores. This stage alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote produced when a haploid egg cell is fertilized by a haploid sperm and each sporophyte cell therefore has a double set of chromosomes, one set from each parent. All land plants, and most multicellular algae, have life cycles in which a multicellular diploid sporophyte phase alternates with a multicellular haploid gametophyte phase. In the seed plants, the largest groups of which are the gymnosperms and flowering plants (angiosperms), the sporophyte phase is more prominent than the gametophyte, and is the familiar green plant with its roots, stem, leaves and cones or flowers. In flowering plants the gametophytes are very reduced in size, and are represented by the germinated pollen and the embryo sac. The sporophyte produces spores (hence t ...
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Palhinhaea Cernua
''Palhinhaea'' is a genus of lycophytes in the family Lycopodiaceae. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is placed in the subfamily Lycopodielloideae. Some sources do not recognize the genus, sinking it into ''Lycopodiella''. ''Palhinhaea'' species are widespread in the tropics and subtropics. Species , the ''Checklist of Ferns and Lycophytes of the World'' recognized the following species: *'' Palhinhaea bradei'' (Nessel) Holub *'' Palhinhaea brevibracteata'' (Alderw.) Holub *'' Palhinhaea camporum'' (B.Øllg. & P.G.Windisch) Holub *''Palhinhaea cernua'' (L.) Vasc. & Franco *'' Palhinhaea cerrojefensis'' B.Øllg. *''Palhinhaea crassifolia'' (Spring) Fraser-Jenk. & Kholia *''Palhinhaea curvata'' (Sw.) Holub *'' Palhinhaea descendens'' (B.Øllg.) Holub *''Palhinhaea divaricata'' B.Øllg. *'' Palhinhaea eichleri'' (Fée) Holub *''Palhinhaea glaucescens'' (C.Presl) Holub *'' Palhinhaea hainanensis'' C.Y.Yang *''Palhinhaea lehmannii'' (Hieron.) Holub ...
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