Hyolith
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Hyolith
Hyoliths are animals with small conical surface, conical animal shell, shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as being basal lophotrochozoans, or even Mollusca, molluscs. Morphology The shell of a hyolith is typically one to four centimeters in length, triangular or elliptical in cross section. Some species have rings or stripes. It comprises two parts: the main conical shell (previously referred to as a ‘conch’) and a cap-like Operculum (animal), operculum. Some also had two curved supports known as ''helens'' They are calcium carbonate, calcareous – probably aragonitic All of these structures grew by marginal accretion. Shell microstructure The orthothecid shell has an internal layer with a microstructure of transverse bundles, and an external layer comprising longitudinal bundles. Helens Some hyoliths ha ...
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Hyoliths02
Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as being basal lophotrochozoans, or even molluscs. Morphology The shell of a hyolith is typically one to four centimeters in length, triangular or elliptical in cross section. Some species have rings or stripes. It comprises two parts: the main conical shell (previously referred to as a ‘conch’) and a cap-like operculum. Some also had two curved supports known as ''helens'' They are calcareous – probably aragonitic All of these structures grew by marginal accretion. Shell microstructure The orthothecid shell has an internal layer with a microstructure of transverse bundles, and an external layer comprising longitudinal bundles. Helens Some hyoliths had helens, long structures that taper as they coil gently in a logarithmic spira ...
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Hyolithida
The Hyolithida are lophophorates, one of the two orders of hyolithid, the other being the Orthothecida The orthothecids are one of the two hyolith orders. Marek diagnoses the order thus: Conchs with a flat or concave ventral surface — opercula with large, flat cardinal processes but without clavicles – tightly sigmoidal, sediment-filled intes .... Most of our knowledge of the hyolithids comes from studies on the Hyolithida. Both orders had an operculum that was not hinged to the conch. However, the Hyolithida are distinct from the Orthothecida in having additional paired, curved, whiskerlike appendages. The Hyolithida were probably bottom feeders living in shallow water, and had tentacules. References Further reading * * Paleozoic invertebrates Prehistoric animal orders Hyolitha {{paleo-protostome-stub ...
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Haplophrentis
''Haplophrentis'' is a genus of tiny shelled hyolithid which lived in the Cambrian Period. Its shell was long and conical, with the open end protected by an operculum, from which two fleshy arms called ''helens'' protruded at the sides. These arms served to elevate the opening of the shells above the sea floor, acting like stilts. Morphology Shell length of ''H. reesi'' reached up to while ''H. carinatus'' reached up to . Juveniles could of course be smaller. It is distinguished from ''Hyolithes'' by the presence of a longitudinal septum on the middle of the inner surface of the top of the shell. Its soft anatomy comprises 12(H. carinatus) to 16 (H. reesi) tentacles attached to a horseshoe-shaped lophophore. A pair of wide structures of uncertain function extend along the length of the conical shell. A larval shell is attached to the shell apex. Affinity The soft anatomy of ''Haplophrentis'' was key to establishing the hyoliths as members of the Lophophorata, the group ...
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Orthothecida
The orthothecids are one of the two hyolith orders. Marek diagnoses the order thus: Conchs with a flat or concave ventral surface — opercula with large, flat cardinal processes but without clavicles – tightly sigmoidal, sediment-filled intestine – helens absent. Sometimes the Circothecidae and Tetrathecidae are split out into a separate order 'Circothecida', which is defined by the bottom surface not being flat, the cardinal processes being pronounced, and a circular rim sometimes showing hints of differentiation into clavicles. Internal taxonomy Marek gives the following diagnoses: * Orthothecidae: Flat or concave bottom surface of conical shell. Transverse aperture. Kouchinsky lists the following taxonomic criteria: * Circothecidae: circular cross-section; same shape and sculpture on upper and lower surfaces. * Turcurthecidae: oval or lenticular cross-section; top and bottom identical * Allathecidae: top side flat or slightly convex; bottom side convex Elsewhere i ...
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Haplophrentis Burgess Shale
''Haplophrentis'' is a genus of tiny shelled hyolithid which lived in the Cambrian Period. Its shell was long and conical, with the open end protected by an operculum, from which two fleshy arms called ''helens'' protruded at the sides. These arms served to elevate the opening of the shells above the sea floor, acting like stilts. Morphology Shell length of ''H. reesi'' reached up to while ''H. carinatus'' reached up to . Juveniles could of course be smaller. It is distinguished from ''Hyolithes'' by the presence of a longitudinal septum on the middle of the inner surface of the top of the shell. Its soft anatomy comprises 12(H. carinatus) to 16 (H. reesi) tentacles attached to a horseshoe-shaped lophophore. A pair of wide structures of uncertain function extend along the length of the conical shell. A larval shell is attached to the shell apex. Affinity The soft anatomy of ''Haplophrentis'' was key to establishing the hyoliths as members of the Lophophorata, the group c ...
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Spence Shale
The Spence Shale is the middle member of the Langston Formation in southeastern Idaho and northeastern Utah. It is exposed in the Bear River Range, the Wasatch Range and the Wellsville Mountains. It is known for its abundant Cambrian trilobites and the preservation of Burgess Shale-type fossils. The type locality is Spence Gulch in southeastern Idaho, near the town of Liberty. It was first described by Charles Doolittle Walcott in 1908. Stratigraphy The Spence Shale spans the '' Albertella'' and '' Glossopleura'' biozones. Fauna Generic list of the fauna of the Spence Shale:Hammersburg, S.R., Hasiotis, S.T., Robison, R.R. 2018. Ichnotaxonomy of the Cambrian Spence Shale Member of the Langston Formation, Wellsville Mountains, Northern Utah, USA. Paleontological Contributions, 20, 1–66. Arthropoda Soft-bodied *''Anomalocaris'' *''Canadaspis'' *'' Caryosyntrips'' *'' Dioxycaris'' *''Hurdia'' *''Isoxys'' *''Leanchoilia'' *'' Meristosoma'' *''Mollisonia'' *''Sidneyia'' * ...
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Lophophore
The lophophore () is a characteristic feeding organ possessed by four major groups of animals: the Brachiopoda, Bryozoa, Hyolitha, and Phoronida, which collectively constitute the protostome group Lophophorata.Introduction to the Lophotrochozoa
– Retrieved 3 May 2010
All lophophores are found in aquatic organisms.


Etymology

''Lophophore'' is derived from the Greek ''lophos'' (crest, tuft) and ''-phore'', ''-phoros'' (φορος) (bearing), a derivative of ''phérein'' (φέρειν) (to bear); thus crest-bearing.


Characteristics

The lophophore can most easily be described as a ring of ted tentacles surrounding the mouth, but it is often horseshoe-shape ...
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Mollusca
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastropod ...
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Lophotrochozoa
Lophotrochozoa (, "crest/wheel animals") is a clade of protostome animals within the Spiralia. The taxon was established as a monophyletic group based on molecular evidence. The clade includes animals like annelids, molluscs, bryozoans, brachiopods, and platyhelminthes. Groups Lophotrochozoa was defined in 1995 as the "last common ancestor of the three traditional lophophorate taxa (brachiopods, bryozoans, and phoronid worms), the mollusks and the annelids, and all of the descendants of that common ancestor". It is a cladistic definition (a node-based name), so the affiliation to Lophotrochozoa of spiralian groups not mentioned directly in the definition depends on the topology of the spiralian tree of life, and in some phylogenetic hypotheses, Lophotrochozoa may even be synonymous to Spiralia. Nemertea and Orthonectida (if not directly considered as part of Annelida) are probably lophotrochozoan phyla; Dicyemida, Gastrotricha, and Platyhelminthes may be lophotrochozoans ...
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Sipunculid
The Sipuncula or Sipunculida (common names sipunculid worms or peanut worms) is a class containing about 162 species of unsegmented marine annelid worms. The name ''Sipuncula'' is from the genus name ''Sipunculus'', and comes from the Latin ''siphunculus'' meaning a "small tube". Sipuncula was once considered a phylum, but was demoted to a class of Annelida, based on recent molecular work. Sipunculans vary in size but most species are under in length. The body is divided into an unsegmented, bulbous trunk and a narrower, anterior section, called the "introvert", which can be retracted into the trunk. The mouth is at the tip of the introvert and is surrounded in most groups by a ring of short tentacles. With no hard parts, the body is flexible and mobile. Although found in a range of habitats throughout the world's oceans, the majority of species live in shallow water habitats, burrowing under the surface of sandy and muddy substrates. Others live under stones, in rock crevic ...
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Journal Of Paleontology
The ''Journal of Paleontology'' is a peer-reviewed scientific journal covering the field of paleontology. It is published by the Paleontological Society. Indexing The ''Journal of Paleontology'' is indexed in: *BIOSIS Previews *Science Citation Index *The Zoological Record *GeoRef __NOTOC__ The GeoRef database is a bibliographic database that indexes scientific literature in the geosciences, including geology. Coverage ranges from 1666 to the present for North American literature, and 1933 to the present for the rest of t ... References Paleontology journals Publications established in 1927 Academic journals published by learned and professional societies Cambridge University Press academic journals Bimonthly journals Paleontological Society {{paleo-journal-stub ...
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Extant Taxon
Neontology is a part of biology that, in contrast to paleontology, deals with living (or, more generally, '' recent'') organisms. It is the study of extant taxa (singular: extant taxon): taxa (such as species, genera and families) with members still alive, as opposed to (all) being extinct. For example: * The moose (''Alces alces'') is an extant species, and the dodo (''Raphus cucullatus'') is an extinct species. * In the group of molluscs known as the cephalopods, there were approximately 600 extant species and 7,500 extinct species. A taxon can be classified as extinct if it is broadly agreed or certified that no members of the group are still alive. Conversely, an extinct taxon can be reclassified as extant if there are new discoveries of living species ("Lazarus species"), or if previously-known extant species are reclassified as members of the taxon. Most biologists, zoologists, and botanists are in practice neontologists, and the term neontologist is used large ...
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