Honey Bee Pheromone
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Honey Bee Pheromone
The pheromones of the honey bee are mixtures of chemical substances released by individual bees into the hive or environment that cause changes in the physiology and behaviour of other bees. Introduction Honey bees (''Apis mellifera'') have one of the most complex pheromonal communication systems found in nature, possessing 15 known glands that produce an array of compounds.Free, John B., ''Pheromones of social bees.'' Ithaca, N.Y.: Comstock, 1987.Blum, M.S. 1992. Honey bee pheromones in The Hive and the Honey Bee, revised edition (Dadant and Sons: Hamilton, Illinois), pages 385–389. These chemical messengers secreted by a queen, drone, worker bee or laying worker bee to elicit a response in other bees. The chemical messages are received by the bee's antenna and other body parts. They are produced as a volatile or non-volatile liquid and transmitted by direct contact as a liquid or vapor. Honey bee pheromones can be grouped into releaser pheromones which temporarily affect the ...
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Pheromone
A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavior of the receiving individuals. There are ''alarm signal, alarm pheromones'', ''food trail pheromones'', ''sex pheromones'', and many others that affect behavior or physiology. Pheromones are used by many organisms, from basic unicellular prokaryotes to complex multicellular eukaryotes. Their use among insects has been particularly well documented. In addition, some vertebrates, plants and ciliates communicate by using pheromones. The ecological functions and evolution of pheromones are a major topic of research in the field of chemical ecology. Background The portmanteau word "pheromone" was coined by Peter Karlson and Martin Lüscher in 1959, based on the Greek φερω ''pheroo'' ('I carry') and ὁρμων ''hormon'' ('stimulating'). P ...
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Swarming
Swarm behaviour, or swarming, is a collective behaviour exhibited by entities, particularly animals, of similar size which aggregate together, perhaps milling about the same spot or perhaps moving ''en masse'' or migrating in some direction. It is a highly interdisciplinary topic. As a term, ''swarming'' is applied particularly to insects, but can also be applied to any other entity or animal that exhibits swarm behaviour. The term '' flocking'' or ''murmuration'' can refer specifically to swarm behaviour in birds, ''herding'' to refer to swarm behaviour in tetrapods, and ''shoaling'' or ''schooling'' to refer to swarm behaviour in fish. Phytoplankton also gather in huge swarms called ''blooms'', although these organisms are algae and are not self-propelled the way animals are. By extension, the term "swarm" is applied also to inanimate entities which exhibit parallel behaviours, as in a robot swarm, an earthquake swarm, or a swarm of stars. From a more abstract point of v ...
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Queen Mandibular Pheromone
Queen mandibular pheromone, or QMP, is a honey bee pheromone produced by the queen and fed to her attendants who share it with the rest of the colony that gives the colony the sense of belonging to the queen. Newly emerged queens produce very little QMP. By the sixth day they are producing enough to attract drones for mating. A laying queen makes twice that amount. Lack of QMP seems to attract robber bees. A study of foraging worker bees has suggested that foraging bees are not attracted to QMP. Chemical composition Chemically, QMP is very diverse, with at least 17 major components and other minor ones. Five of these compounds are: 9-oxo-2-decenoic acid (9ODA), ''cis''- and ''trans''-9-hydroxydec-2-enoic acid (9HDA), methyl ''p''-hydroxybenzoate (HOB) and 4-hydroxy-3-methoxyphenylethanol (HVA). Drones QMP functions as a sex pheromone for drones, attracting males to an unmated queen. 9ODA specifically is known to attract drones over long distances, and its combination with 9HDA ...
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Tergite
A ''tergum'' (Latin for "the back"; plural ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'margin'. A given tergum may be divided into hardened plates or sclerites commonly referred to as tergites. In a thoracic segment, for example, the tergum may be divided into an anterior notum and a posterior scutellum. Lateral extensions of a tergite are known as paranota (Greek for "alongside the back") or ''carinae'' (Latin for "keel"), exemplified by the flat-backed millipedes of the order Polydesmida. Kinorhynchs have tergal and sternal plates too, though seemingly not homologous with those of arthropods. Tergo-tergal is a stridulatory mechanism in which fine spines of the abdominal tergites are rubbed together to produce sound. This process is known as abdominal telescoping. Examples File:Andrena spiraeana abdomen.jpg , Abdominal t ...
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Geranic Acid
Geranic acid, or 3,7-dimethyl-2,6-octadienoic acid, is a pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ... used by some organisms. It is a double bond isomer of nerolic acid. Pharmacology Choline geranate (also described as Choline And Geranic acid, or CAGE) has been developed as a novel biocompatible antiseptic material capable of penetrating skin and aiding the transdermal delivery of co-administered antibiotics. The antibacterial properties of CAGE were analyzed against 24 and 72 hour old biofilms of 11 clinically isolated ESKAPE pathogens (defined as Enterococcus faecium, Staphylococcus aureus, Klebsiella pneumoniae, Acinetobacter baumanii, Pseudomonas aeruginosa, and Enterobacter sp, respectively), including multidrug resistant (MDR) isolates. CAGE was observ ...
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Citral
Citral is an acyclic monoterpene aldehyde, and being a monoterpene, it is made of two isoprene units. Citral is a collective term which covers two geometric isomers that have their own separate names; the ''E''-isomer is named geranial (''trans''-citral) or citral A. The ''Z''-isomer is named neral (''cis''-citral) or citral B. These stereoisomers occur as a mixture, not necessarily racemic; e.g. in essential oil of Australian ginger, the neral to geranial ratio is 0.61. Occurrence Citral is present in the oils of several plants, including lemon myrtle (90–98%), '' Litsea citrata'' (90%), ''Litsea cubeba'' (70–85%), lemongrass (65–85%), lemon tea-tree (70–80%), ''Ocimum gratissimum'' (66.5%), '' Lindera citriodora'' (about 65%), '' Calypranthes parriculata'' (about 62%), petitgrain (36%), lemon verbena (30–35%), lemon ironbark (26%), lemon balm (11%), lime (6–9%), lemon (2–5%), and orange. Further, in the lipid fraction (essential oil) of Australian ginger (51- ...
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Nerolic Acid
Nerolic acid, also known as (''Z'')-3,7-Dimethyl-2,6-octadienoic acid, is an organically-derived chemical. In nature Nerolic acid is found in the Nasonov scent gland of honey-bees along with geraniol, geranic acid, citral, farnesol, and nerol. Of these, nerolic acid, geraniol, and farnesol are present in the highest proportions.. It is one of the five compounds that are part of the essential oils of ''Myrcia Ovata''. It is a compound also found in great quantity in ''Myrcia lundiana'' and it possess antifungal properties against pathogens such as ''Fusarium solani,'' and ''Lasiodiplodia theobromae''. Both ''Myrcia Ovata'' and ''Myrcia lundiana'' are part of the Myrtaceae family plant, both containing a certain percentage of Nerolic acid compound. In addition to this, Nerolic acid is also a principal chemical compound of essential oils of lemongrass ''Cymbopogon'', also known as lemongrass, barbed wire grass, silky heads, Cochin grass, Malabar grass, oily heads, cit ...
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Geraniol
Geraniol is a monoterpenoid and an alcohol. It is the primary component of citronella oil and is a primary component of rose oil, palmarosa oil. It is a colorless oil, although commercial samples can appear yellow. It has low solubility in water, but it is soluble in common organic solvents. The functional group derived from geraniol (in essence, geraniol lacking the terminal −OH) is called geranyl. Uses and occurrence In addition to rose oil, palmarosa oil, and citronella oil, it also occurs in small quantities in geranium, lemon, and many other essential oils. With a rose-like scent, it is commonly used in perfumes. It is used in flavors such as peach, raspberry, grapefruit, red apple, plum, lime, orange, lemon, watermelon, pineapple, and blueberry. Geraniol is produced by the scent glands of honeybees to mark nectar-bearing flowers and locate the entrances to their hives. It is also commonly used as an insect repellent, especially for mosquitoes. The scent of geraniol ...
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Nasonov Pheromone
The Nasonov (alternatively, Nasanov) pheromone is released by worker bees to orient returning forager bees back to the colony. To broadcast this scent, bees raise their abdomens, which contain the Nasonov glands, and fan their wings vigorously. Nasonov includes a number of different terpenoids including geraniol, nerolic acid, citral and geranic acid. Bees use these to find the entrance to their colony or hive, and they release them on flowers so other bees know which flowers have nectar. Once the foraging bee leaves the nest it uses its sense from special sensing cells on the antennae to locate and distinguish forage plants, which each give off a unique blend of odour chemicals. When a beekeeper lifts out frames from a hive they disturb the balance of smells within the hive. It can take up to 48 hours for the colony to re-establish its scent equilibrium. A synthetically produced Nasonov pheromone can be used to attract a honey bee swarm to an unoccupied hive or a swarm-ca ...
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Ethyl Oleate
Ethyl oleate is a fatty acid ester formed by the condensation of oleic acid and ethanol. It is a colorless oil although degraded samples can appear yellow. Use and occurrence Additive Ethyl oleate is used by compounding pharmacies as a vehicle for intramuscular drug delivery, in some cases to prepare the daily doses of progesterone in support of pregnancy. Studies that document the safe use of ethyl oleate in pregnancy for both the mother and the fetus have never been performed. It is regulated as a food additive in the U.S. by the Food and Drug Administration. Ethyl oleate is used as a solvent for pharmaceutical drug preparations involving lipophilic substances such as steroids. It also finds use as a lubricant and a plasticizer. Louis Bouveault used ethyl oleate to demonstrate Bouveault–Blanc reduction, producing oleyl alcohol and ethanol, a method which was subsequently refined and published in ''Organic Syntheses''. Occurrence Ethyl oleate has been identified as a pri ...
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Tarsal Gland
Meibomian glands (also called tarsal glands, palpebral glands, and tarsoconjunctival glands) are sebaceous glands along the rims of the eyelid inside the tarsal plate. They produce meibum, an oily substance that prevents evaporation of the eye's tear film. Meibum prevents tears from spilling onto the cheek, traps them between the oiled edge and the eyeball, and makes the closed lids airtight. There are about 25 such glands on the upper eyelid, and 20 on the lower eyelid. Dysfunctional meibomian glands is believed to be the most often cause of dry eyes. They are also the cause of posterior blepharitis. History The glands were mentioned by Galen in 200 AD and were described in more detail by Heinrich Meibom (1638–1700), a German physician, in his work ''De Vasis Palpebrarum Novis Epistola'' in 1666. This work included a drawing with the basic characteristics of the glands. Anatomy Although the upper lid have greater number and volume of meibomian glands than the lower li ...
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