Histotrophy
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Histotrophy
Histotrophy is a form of matrotrophy exhibited by some live-bearing sharks and rays, in which the developing embryo receives additional nutrition from its mother in the form of uterine secretions, known as histotroph (or "uterine milk"). It is one of the three major modes of elasmobranch reproduction encompassed by " aplacental viviparity", and can be contrasted with yolk-sac viviparity (in which the embryo is solely sustained by yolk) and oophagy (in which the embryo feeds on ova). There are two categories of histotrophy: *In mucoid or limited histotrophy, the developing embryo ingests uterine mucus or histotroph as a supplement to the energy supplies provided by its yolk sac. This form of histotrophy is known to occur in the dogfish sharks (Squaliformes) and the electric rays (Torpediniformes), and may be more widespread. *In lipid histotrophy, the developing embryo is supplied with protein and lipid-enriched histotroph through specialized finger-like structures known as trophone ...
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Matrotrophy
Matrotrophy is a form of maternal care during organism development, associated with live birth (viviparity), in which the embryo of an animal or flowering plant is supplied with additional nutrition from the mother (e.g. through a placenta). This can be contrasted with lecithotrophy, in which the only source of nutrition for the embryo is yolk originally contained within its egg. Vegetal matrotrophy In plants, matrotrophy is considered a critical evolutionary development preceding the origin of embryophytes and therefore essential to the evolution of land plants. Matrotrophy is facilitated by cytological and ultrastructural modifications on one or both sides of the generational junction, a region called the placenta. Specialization of the placental cells pertains further to their cytological and ultrastructural characteristics: the cytoplasm is often dense and rich in lipids, the vacuole is typically reduced but large in ''Sphagnum'', the endoplasmic reticulum extensive, mitochon ...
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Yolk Sac
The yolk sac is a membranous sac attached to an embryo, formed by cells of the hypoblast layer of the bilaminar embryonic disc. This is alternatively called the umbilical vesicle by the Terminologia Embryologica (TE), though ''yolk sac'' is far more widely used. In humans, the yolk sac is important in early embryonic blood supply, and much of it is incorporated into the primordial gut during the fourth week of embryonic development. In humans The yolk sac is the first element seen within the gestational sac during pregnancy, usually at 3 days gestation. The yolk sac is situated on the front (ventral) part of the embryo; it is lined by extra-embryonic endoderm, outside of which is a layer of extra-embryonic mesenchyme, derived from the epiblast. Blood is conveyed to the wall of the yolk sac by the primitive aorta and after circulating through a wide-meshed capillary plexus, is returned by the vitelline veins to the tubular heart of the embryo. This constitutes the vitell ...
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Embryology
Embryology (from Greek ἔμβρυον, ''embryon'', "the unborn, embryo"; and -λογία, '' -logia'') is the branch of animal biology that studies the prenatal development of gametes (sex cells), fertilization, and development of embryos and fetuses. Additionally, embryology encompasses the study of congenital disorders that occur before birth, known as teratology. Early embryology was proposed by Marcello Malpighi, and known as preformationism, the theory that organisms develop from pre-existing miniature versions of themselves. Aristotle proposed the theory that is now accepted, epigenesis. Epigenesis is the idea that organisms develop from seed or egg in a sequence of steps. Modern embryology, developed from the work of Karl Ernst von Baer, though accurate observations had been made in Italy by anatomists such as Aldrovandi and Leonardo da Vinci in the Renaissance. Comparative embryology Preformationism and epigenesis As recently as the 18th century, the prevailin ...
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Myliobatiformes
Myliobatiformes () is one of the four orders of batoids, cartilaginous fishes related to sharks. They were formerly included in the order Rajiformes, but more recent phylogenetic studies have shown the myliobatiforms to be a monophyletic group, and its more derived members evolved their highly flattened shapes independently of the skates. Classification Nelson's ''Fishes of the World'' arranges the Myliobatiformes as: *Suborder Platyrhinoidei **Family Platyrhinidae (thornbacks) *Suborder Zanobatoidei **Family Zanobatidae (panrays) *Suborder Myliobatoidei ( stingrays) **Superfamily Hexatrygonoidea ***Family Hexatrygonidae (sixgill stingray) **Superfamily Urolophoidea ***Family Plesiobatidae (deepwater stingray) ***Family Urolophidae (round stingrays) **Superfamily Urotrygonoidea ***Family Urotrygonidae (American round stingrays) **Superfamily Dasyatoidea ***Family Dasyatidae (whiptail stingrays) ***Family Potamotrygonidae (river stingrays) ***Family Gymnuridae The bu ...
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Orders Of Magnitude
An order of magnitude is an approximation of the logarithm of a value relative to some contextually understood reference value, usually 10, interpreted as the base of the logarithm and the representative of values of magnitude one. Logarithmic distributions are common in nature and considering the order of magnitude of values sampled from such a distribution can be more intuitive. When the reference value is 10, the order of magnitude can be understood as the number of digits in the base-10 representation of the value. Similarly, if the reference value is one of some powers of 2, since computers store data in a binary format, the magnitude can be understood in terms of the amount of computer memory needed to store that value. Differences in order of magnitude can be measured on a base-10 logarithmic scale in “decades” (i.e., factors of ten). Examples of numbers of different magnitudes can be found at Orders of magnitude (numbers). Definition Generally, the order of magnitude ...
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Lipid
Lipids are a broad group of naturally-occurring molecules which includes fats, waxes, sterols, fat-soluble vitamins (such as vitamins A, D, E and K), monoglycerides, diglycerides, phospholipids, and others. The functions of lipids include storing energy, signaling, and acting as structural components of cell membranes. Lipids have applications in the cosmetic and food industries, and in nanotechnology. Lipids may be broadly defined as hydrophobic or amphiphilic small molecules; the amphiphilic nature of some lipids allows them to form structures such as vesicles, multilamellar/unilamellar liposomes, or membranes in an aqueous environment. Biological lipids originate entirely or in part from two distinct types of biochemical subunits or "building-blocks": ketoacyl and isoprene groups. Using this approach, lipids may be divided into eight categories: fatty acyls, glycerolipids, glycerophospholipids, sphingolipids, saccharolipids, and polyketides (derived from condensati ...
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Protein
Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, responding to stimuli, providing structure to cells and organisms, and transporting molecules from one location to another. Proteins differ from one another primarily in their sequence of amino acids, which is dictated by the nucleotide sequence of their genes, and which usually results in protein folding into a specific 3D structure that determines its activity. A linear chain of amino acid residues is called a polypeptide. A protein contains at least one long polypeptide. Short polypeptides, containing less than 20–30 residues, are rarely considered to be proteins and are commonly called peptides. The individual amino acid residues are bonded together by peptide bonds and adjacent amino acid residues. The sequence of amino acid residue ...
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Electric Ray
The electric rays are a group of rays, flattened cartilaginous fish with enlarged pectoral fins, composing the order Torpediniformes . They are known for being capable of producing an electric discharge, ranging from 8 to 220 volts, depending on species, used to stun prey and for defense. There are 69 species in four families. Perhaps the best known members are those of the genus ''Torpedo''. The torpedo undersea weapon is named after it. The name comes from the Latin , 'to be stiffened or paralyzed', from the effect on someone who touches the fish. Description Electric rays have a rounded pectoral disc with two moderately large rounded-angular (not pointed or hooked) dorsal fins (reduced in some Narcinidae), and a stout muscular tail with a well-developed caudal fin. The body is thick and flabby, with soft loose skin with no dermal denticles or thorns. A pair of kidney-shaped electric organs are at the base of the pectoral fins. The snout is broad, large in the Narcinidae, b ...
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Squaliformes
The Squaliformes are an order (biology), order of sharks that includes about 126 species in seven families. Members of the order have two dorsal fins, which usually possess spines, they usually have a sharp head, no anal fin or nictitating membrane, and five to seven gill slits. In most other respects, however, they are quite variable in form and size. Most species of the squaliform order live in saltwater or brackish water. They are found worldwide, from northern to tropical waters, and from shallow coastal seas to the open ocean. All members of the family Eptomeridae and Dalatiidae and ''Zameus squamulosus'' possess photophores, luminous organs, and exhibit intrinsic bioluminescence. Bioluminescence evolved once in Squaliformes, approximately 111–153 million years ago, and helped the Squaliformes radiate and adapt to the deep sea. The common ancestor of Dalatiidae, Etmopteridae, Somniosidae, and Oxynotidae possessed a luminous organ and used bioluminescence for camouf ...
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Mucus
Mucus ( ) is a slippery aqueous secretion produced by, and covering, mucous membranes. It is typically produced from cells found in mucous glands, although it may also originate from mixed glands, which contain both serous and mucous cells. It is a viscous colloid containing inorganic salts, antimicrobial enzymes (such as lysozymes), immunoglobulins (especially IgA), and glycoproteins such as lactoferrin and mucins, which are produced by goblet cells in the mucous membranes and submucosal glands. Mucus serves to protect epithelial cells in the linings of the respiratory, digestive, and urogenital systems, and structures in the visual and auditory systems from pathogenic fungi, bacteria and viruses. Most of the mucus in the body is produced in the gastrointestinal tract. Amphibians, fish, snails, slugs, and some other invertebrates also produce external mucus from their epidermis as protection against pathogens, and to help in movement and is also produced in fish to line the ...
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Shark
Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimorpha (or Selachii) and are the sister group to the rays. However, the term "shark" has also been used to refer to all extinct members of Chondrichthyes with a shark-like morphology, such as hybodonts and xenacanths. The oldest modern sharks are known from the Early Jurassic. They range in size from the small dwarf lanternshark (''Etmopterus perryi''), a deep sea species that is only in length, to the whale shark (''Rhincodon typus''), the largest fish in the world, which reaches approximately in length. Sharks are found in all seas and are common to depths up to . They generally do not live in freshwater, although there are a few known exceptions, such as the bull shark and the river shark, which can be found in both seawater and fresh ...
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Ovum
The egg cell, or ovum (plural ova), is the female reproductive cell, or gamete, in most anisogamous organisms (organisms that reproduce sexually with a larger, female gamete and a smaller, male one). The term is used when the female gamete is not capable of movement (non-motile). If the male gamete (sperm) is capable of movement, the type of sexual reproduction is also classified as oogamous. A nonmotile female gamete formed in the oogonium of some algae, fungi, oomycetes, or bryophytes is an oosphere. When fertilized the oosphere becomes the oospore. When egg and sperm fuse during fertilisation, a diploid cell (the zygote) is formed, which rapidly grows into a new organism. History While the non-mammalian animal egg was obvious, the doctrine ''ex ovo omne vivum'' ("every living nimal comes froman egg"), associated with William Harvey (1578–1657), was a rejection of spontaneous generation and preformationism as well as a bold assumption that mammals also reproduced via ...
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