Harvester Termites
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Harvester Termites
The harvester termites (from Greek ὁδός (hodós), travelling; Latin '' termes'', woodworm) are an ancient, Old World family of termites, the Hodotermitidae. They are distinguished by the serrated inner edge of their mandibles, and their functional compound eyes which are present in all castes. They forage for grass at night and during daylight hours, and pigmented workers are often observed outside the nest. Their range includes the deserts and savannas of Africa, the Middle East, and Southwest Asia. Their English name refers to their habit of collecting grass, which is not unique to the family, though. General The family consists of three extant genera and some 18 or 19 species. ''Anacanthotermes'' is found in deserts and semideserts of North Africa, the Middle East, and Southwest Asia, including Baluchistan and southern India. ''Hodotermes'' has a vast range from Palaearctic North Africa, through the East African savannas to the karroid regions of southern Africa. ''Microho ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Old World
The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by their inhabitants as comprising the entire world, with the "New World", a term for the newly encountered lands of the Western Hemisphere, particularly the Americas. Etymology In the context of archaeology and world history, the term "Old World" includes those parts of the world which were in (indirect) cultural contact from the Bronze Age onwards, resulting in the parallel development of the early civilizations, mostly in the temperate zone between roughly the 45th and 25th parallels north, in the area of the Mediterranean, including North Africa. It also included Mesopotamia, the Persian plateau, the Indian subcontinent, China, and parts of Sub-Saharan Africa. These regions were connected via the Silk Road trade route, and they have a p ...
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Pteronia
''Pteronia'' ("resin daisies") is a genus of evergreen, woody perennial plants assigned to the family Asteraceae with currently 76 described species. Like in almost all Asteraceae, the individual flowers are 5-merous, small and clustered in typical heads, surrounded by an involucre of bracts. In ''Pteronia'', the centre of the head is taken by relatively few, yellow, disc florets, while a ring of ligulate florets is absent. These florets sit on a common base (or receptacle). Taxonomy A species of Gombos was first described and assigned to the new genus ''Pteronia'' by the famous Swedish naturalist Carl Linnaeus in the second edition of his groundbreaking Species Plantarum, the starting point of modern botanical nomenclature, that was published in 1763. He had not seen living plants or dried herbarium specimens, but based his description on an etching made by the English botanist Leonard Plukenet in 1700. This etching probably represents '' Pteronia camphorata'', which has b ...
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Alate
Alate (Latin ''ālātus'', from ''āla'' (“wing”)) is an adjective and noun used in entomology and botany to refer to something that has wings or winglike structures. In entomology In entomology, "alate" usually refers to the winged form of a social insect, especially ants or termites, though can also be applied to aphids and some thrips. An alate is a winged reproductive caste from a social insect colony in its winged form. Alate females are typically those destined to become gynes (queens), whereas alate males are occasionally referred to as "drones" (or "kings", in the case of termites). Their common behavioural function is starting a new colony, to expand their mother colonies etc. Colonies of termites and ants produce alates. It is a flight-based form of reproductive technique. In a termite colony, alates (winged males and winged females) disperse in a specific period or a month. Male and female pair to each other during flight, shed their wings, and start a new colo ...
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Microhodotermes Viator
''Microhodotermes'' is a genus of southern African harvester termites in the Hodotermitidae. As with harvester termites in general, they have serrated inner edges to their mandibles, and all castes have functional compound eyes. Species of this genus are desert specialists of the Namib, Kalahari and Karoo, where their ranges overlap with ''Hodotermes''. They forage at night and during daylight hours, and their pigmented workers are often observed outside the nest. The workers of '' M. viator'' collect mostly woody material, with ''Pteronia'' and vygie species being favoured. Colonies of ''M. viator'' produce initially small, conical mounds on soil with sufficient clay content. These are speculated to cause the formation of increasingly large heuweltjies. Widespread foraging and burrowing activity of aardvark The aardvark ( ; ''Orycteropus afer'') is a medium-sized, burrowing, nocturnal mammal native to Africa. It is the only living species of the order Tubulidentata, ...
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Kalotermitidae
Kalotermitidae is a family of termites, commonly known as drywood termites. Kalotermitidae includes 21 genera and 419 species. The family has a cosmopolitan circumtropical distribution, and is found in functionally arid environments. Biology The Kalotermitidae are "primitive" in morphology, nesting behavior, and social organization. Unlike other termite species, they have no need to make contact with soil and live exclusively within excavations in wood, lacking elaborate nesting architecture. Drywood termites have an adaptive mechanism for conserving water. Undigested matter in the alimentary canal passes through specialized rectal glands in the hindgut. These glands reabsorb water from the feces. They can tolerate dry conditions for long periods of time, receiving all of the moisture they need from the wood they live in and consume. Their mandibles are also fortified with zinc, as an adaptation to the mechanically difficult food source of dry wood. Their diet of dry wood makes ...
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Isoptera
Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattodea (along with cockroaches). Termites were once classified in a separate order from cockroaches, but recent phylogenetic studies indicate that they evolved from cockroaches, as they are deeply nested within the group, and the sister group to wood eating cockroaches of the genus ''Cryptocercus''. Previous estimates suggested the divergence took place during the Jurassic or Triassic. More recent estimates suggest that they have an origin during the Late Jurassic, with the first fossil records in the Early Cretaceous. About 3,106 species are currently described, with a few hundred more left to be described. Although these insects are often called "white ants", they are not ants, and are not closely related to ants. Like ants and some bees an ...
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Plesiomorph
In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and synapomorphy, all mean a trait shared between species because they share an ancestral species. Apomorphic and synapomorphic characteristics convey much information about evolutionary clades and can be used to define taxa. However, plesiomorphic and symplesiomorphic characteristics cannot. The term ''symplesiomorphy'' was introduced in 1950 by German entomologist Willi Hennig. Examples A backbone is a plesiomorphic trait shared by birds and mammals, and does not help in placing an animal in one or the other of these two clades. Birds and mammals share this trait because both clades are descended from the same far distant ancestor. Other clades, e.g. snakes, lizards, turtles, fish, frogs, all have backbones and none are either birds nor ...
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Archotermopsidae
Archotermopsidae is a family of termites in the order Blattodea, known as dampwood termites, formerly included within the family Termopsidae. They constitute a small and rather primitive family with five extant genera and 13–20 living species. They may be a nuisance, but compared to the drywood termites (Kalotermitidae), usually do not cause extensive damage to buildings or other man-made structures. As their name implies, they eat wood that is not dried out, perhaps even rotting, and consequently of little use to humans. Taxonomy In 2009, the five extant genera from the family Termopsidae ('' Archotermopsis'', '' Hodotermopsis'', '' Porotermes'', '' Stolotermes'', and ''Zootermopsis'') were moved to a newly created family, Archotermopsidae (''Zootermopsis'' had previously been treated as part of the family Hodotermitidae), so that the family Termopsidae now includes only fossil taxa: ''Asiatermes'', ''Huaxiatermes'', and ''Mesotermopsis'' (Early Cretaceous of China); ''Cretate ...
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Karoo
The Karoo ( ; from the Afrikaans borrowing of the South Khoekhoe !Orakobab or Khoemana word ''ǃ’Aukarob'' "Hardveld") is a semi-desert natural region of South Africa. No exact definition of what constitutes the Karoo is available, so its extent is also not precisely defined. The Karoo is partly defined by its topography, geology and climate, and above all, its low rainfall, arid air, cloudless skies, and extremes of heat and cold.Potgieter, D.J. & du Plessis, T.C. (1972) ''Standard Encyclopaedia of Southern Africa''. Vol. 6. pp. 306–307. Nasou, Cape Town.''Reader’s Digest Illustrated Guide to Southern Africa''. (5th Ed. 1993). pp. 78–89. Reader’s Digest Association of South Africa Pty. Ltd., Cape Town. The Karoo also hosted a well-preserved ecosystem hundreds of million years ago which is now represented by many fossils. The ǃ’Aukarob formed an almost impenetrable barrier to the interior from Cape Town, and the early adventurers, explorers, hunters, and travelers o ...
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Kalahari Desert
The Kalahari Desert is a large semi-arid sandy savanna in Southern Africa extending for , covering much of Botswana, and parts of Namibia and South Africa. It is not to be confused with the Angolan, Namibian, and South African Namib coastal desert, whose name is of Khoekhoegowab origin and means "vast place". Etymology ''Kalahari'' is derived from the Tswana word ''Kgala'', meaning "the great thirst", or ''Kgalagadi'', meaning "a waterless place"; the Kalahari has vast areas covered by red sand without any permanent surface water. History The Kalahari Desert was not always a dry desert. The fossil flora and fauna from Gcwihaba Cave in Botswana indicates that the region was much wetter and cooler at least from 30 to 11 thousand BP (before present) especially after 17,500 BP. Geography Drainage of the desert is by dry black valleys, seasonally inundated pans and the large salt pans of the Makgadikgadi Pan in Botswana and Etosha Pan in Namibia. The only permanent river, ...
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Namib Desert
The Namib ( ; pt, Namibe) is a coastal desert in Southern Africa. The name is of Khoekhoegowab origin and means "vast place". According to the broadest definition, the Namib stretches for more than along the Atlantic coasts of Angola, Namibia, and South Africa, extending southward from the Carunjamba River in Angola, through Namibia and to the Olifants River (Western Cape), Olifants River in Western Cape, South Africa. The Namib's northernmost portion, which extends from the Angola-Namibia border, is known as Moçâmedes Desert, while its southern portion approaches the neighboring Kalahari Desert. From the Atlantic coast eastward, the Namib gradually ascends in elevation, reaching up to inland to the foot of the Great Escarpment, Southern Africa, Great Escarpment. Annual precipitation ranges from in the most arid regions to at the escarpment, making the Namib the only true desert in southern Africa. Having endured Desert climate, arid or Semi-arid climate, semi-arid cond ...
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