Gnathochilarium
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Gnathochilarium
In arthropods, the maxillae (singular maxilla) are paired structures present on the head as mouthparts in members of the clade Mandibulata, used for tasting and manipulating food. Embryologically, the maxillae are derived from the 4th and 5th segment of the head and the maxillary palps; segmented appendages extending from the base of the maxilla represent the former leg of those respective segments. In most cases, two pairs of maxillae are present and in different arthropod groups the two pairs of maxillae have been variously modified. In crustaceans, the first pair are called maxillulae (singular maxillula). Modified coxae at the base of the pedipalps in spiders are also called "maxillae", although they are not homologous with mandibulate maxillae. Myriapoda Millipedes In millipedes, the second maxillae have been lost, reducing the mouthparts to only the first maxillae which have fused together to form a gnathochilarium, acting as a lower lip to the buccal cavity and the man ...
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Millipede
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from the Latin for "thousand feet", no species was known to have 1,000 or more until the discovery of ''Eumillipes persephone'', which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures. Most millipedes are slow-moving detritivores, eating decaying leaves and other dead plant matter. Some eat fungi or ...
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Arthropod Mouthparts
The mouthparts of arthropods have evolved into a number of forms, each adapted to a different style or mode of feeding. Most mouthparts represent modified, paired appendages, which in ancestral forms would have appeared more like legs than mouthparts. In general, arthropods have mouthparts for cutting, chewing, piercing, sucking, shredding, siphoning, and filtering. This article outlines the basic elements of four arthropod groups: insects, myriapods, crustaceans and chelicerates. Insects are used as the model, with the novel mouthparts of the other groups introduced in turn. Insects are not, however, the ancestral form of the other arthropods discussed here. Insects Insect mouthparts exhibit a range of forms. The earliest insects had chewing mouthparts. Specialisation includes mouthparts modified for siphoning, piercing, sucking and sponging. These modifications have evolved a number of times. For example, mosquitoes (which are flies) and aphids (which are bugs) both ...
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General Malacostracan En
A general officer is an officer of high rank in the armies, and in some nations' air forces, space forces, and marines or naval infantry. In some usages the term "general officer" refers to a rank above colonel."general, adj. and n.". OED Online. March 2021. Oxford University Press. https://www.oed.com/view/Entry/77489?rskey=dCKrg4&result=1 (accessed May 11, 2021) The term ''general'' is used in two ways: as the generic title for all grades of general officer and as a specific rank. It originates in the 16th century, as a shortening of ''captain general'', which rank was taken from Middle French ''capitaine général''. The adjective ''general'' had been affixed to officer designations since the late medieval period to indicate relative superiority or an extended jurisdiction. Today, the title of ''general'' is known in some countries as a four-star rank. However, different countries use different systems of stars or other insignia for senior ranks. It has a NATO rank scal ...
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Sternite
The sternum (pl. "sterna") is the ventral portion of a segment of an arthropod thorax or abdomen. In insects, the sterna are usually single, large sclerites, and external. However, they can sometimes be divided in two or more, in which case the subunits are called sternites, and may also be modified on the terminal abdominal segments so as to form part of the functional genitalia, in which case they are frequently reduced in size and development, and may become internalized and/or membranous. For a detailed explanation of the terminology, see Kinorhynchs have tergal and sternal plates too, though seemingly not homologous with those of arthropods.Sørensen, M. V. et al. Phylogeny of Kinorhyncha based on morphology and two molecular loci. PLoS One 10, 1–33 (2015). Ventrites are externally visible sternites. Usually the first sternite is covered up, so that vertrite numbers do not correspond to sternid numbers. The term is also used in other arthropod groups such as crustaceans ...
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Arthropod Anatomy
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior organs; it has an open circulatory system. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. Their nervous system is "ladder-lik ...
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Nymph (biology)
In biology, a nymph is the immature form of some invertebrates, particularly insects, which undergoes gradual metamorphosis (hemimetabolism) before reaching its adult stage. Unlike a typical larva, a nymph's overall form already resembles that of the adult, except for a lack of wings (in winged species). In addition, while a nymph moults, it never enters a pupal stage. Instead, the final moult results in an adult insect. Nymphs undergo multiple stages of development called instars. This is the case, for example, in Orthoptera (crickets, grasshoppers and locusts), Hemiptera (cicadas, shield bugs, whiteflies, aphids, leafhoppers, froghoppers, treehoppers etc.), mayflies, termites, cockroaches, mantises, stoneflies and Odonata (dragonflies and damselflies). Nymphs of aquatic insects, as in the Odonata, Ephemeroptera, and Plecoptera, are also called naiads, an Ancient Greek name for mythological water nymphs. Usage of the term 'naiad' is no longer popular among entomologists, ...
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Odonata
Odonata is an order of flying insects that includes the dragonflies and damselflies. Members of the group first appeared during the Triassic, though members of their total group, Odonatoptera, first appeared in Late Carboniferous. The two common groups are distinguished with dragonflies, placed in the suborder Epiprocta, usually being larger, with eyes together and wings up or out at rest, while damselflies, suborder Zygoptera, are usually smaller with eyes placed apart and wings along body at rest. All Odonata have aquatic larvae called naiads (nymphs), and all of them, larvae and adults, are carnivorous. The adults can land, but rarely walk. Their legs are specialised for catching prey. They are almost entirely insectivorous. Etymology and terminology Fabricius coined the term ''Odonata'' in 1793 from the Ancient Greek ( Ionic form of ) 'tooth'. One hypothesis is that it was because their maxillae are notably toothed. Most insects also have toothed mandibles. The wo ...
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Lepidopterans
Lepidoptera ( ) is an order of insects that includes butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 percent of the total described species of living organisms. It is one of the most widespread and widely recognizable insect orders in the world. The Lepidoptera show many variations of the basic body structure that have evolved to gain advantages in lifestyle and distribution. Recent estimates suggest the order may have more species than earlier thought, and is among the four most speciose orders, along with the Hymenoptera, Diptera, and Coleoptera. Lepidopteran species are characterized by more than three derived features. The most apparent is the presence of scales that cover the bodies, wings, and a proboscis. The scales are modified, flattened "hairs", and give butterflies and moths their wide variety of colors and patterns. Almost all species have some form of membrano ...
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Hemiptera
Hemiptera (; ) is an order (biology), order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, Reduviidae, assassin bugs, Cimex, bed bugs, and shield bugs. They range in size from to around , and share a common arrangement of piercing-sucking Insect mouthparts, mouthparts. The name "true bugs" is often limited to the suborder Heteroptera. Entomologists reserve the term ''bug'' for Hemiptera or Heteroptera,Gilbert Waldbauer. ''The Handy Bug Answer Book.'' Visible Ink, 1998p. 1. which does not include other arthropods or insects of other orders such as Ant, ants, Bee, bees, Beetle, beetles, or Butterfly, butterflies. In some variations of English, all Terrestrial animal, terrestrial arthropods (including non-insect arachnids, and myriapods) also fall under the Colloquialism, colloquial understanding of ''bug''. Many insects with "bug" in their common name, especially in American English, belo ...
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Insect Mouthparts
Insects have mouthparts that may vary greatly across insect species, as they are adapted to particular modes of feeding. The earliest insects had chewing mouthparts. Most specialisation of mouthparts are for piercing and sucking, and this mode of feeding has evolved a number of times idependently. For example, mosquitoes and aphids (which are true bugs) both pierce and suck, however female mosquitoes feed on animal blood whereas aphids feed on plant fluids. Evolution Like most external features of arthropods, the mouthparts of Hexapoda are highly derived. Insect mouthparts show a multitude of different functional mechanisms across the wide diversity of insect species. It is common for significant homology to be conserved, with matching structures forming from matching primordia, and having the same evolutionary origin. However, even if structures are almost physically and functionally identical, they may not be homologous; their analogous functions and appearance might be the pr ...
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Thysanura
Thysanura is the now deprecated name of what was, for over a century, recognised as an order in the class Insecta. The two constituent groups within the former order, the Archaeognatha (jumping bristletails) and the Zygentoma (silverfish and firebrats), share several characteristics, such as of having three long caudal filaments, the lateral ones being the cerci, while the one between (telson) is a medial cerciform appendage, specifically an epiproct. They are also both wingless, and have bodies covered with fine scales, rather like the scales of the practically unrelated Lepidoptera. In the late 20th century, it was recognized that the two suborders were not sister taxa, therefore Thysanura was paraphyletic, and the two suborders were each raised to the status of an independent monophyletic order, with Archaeognatha sister taxon to the Dicondylia, including the Zygentoma. Although the group Thysanura is no longer recognized, the name still appears in some published material. An ...
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Archaeognatha
The Archaeognatha are an order of apterygotes, known by various common names such as jumping bristletails. Among extant insect taxa they are some of the most evolutionarily primitive; they appeared in the Middle Devonian period at about the same time as the arachnids. Specimens that closely resemble extant species have been found as both body and trace fossils (the latter including body imprints and trackways) in strata from the remainder of the Paleozoic Era and more recent periods. For historical reasons an alternative name for the order is Microcoryphia. Until the late 20th century the suborders Zygentoma and Archaeognatha comprised the order Thysanura; both orders possess three-pronged tails comprising two lateral cerci and a medial epiproct or ''appendix dorsalis''. Of the three organs, the appendix dorsalis is considerably longer than the two cerci; in this the Archaeognatha differ from the Zygentoma, in which the three organs are subequal in length. In the late 20th cen ...
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