Escape Reflex
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Escape Reflex
Escape reflex, or escape behavior, is any kind of escape response found in an animal when it is presented with an unwanted stimulus. It is a simple reflectory reaction in response to stimuli indicative of danger, that initiates an escape motion of an animal. The escape response has been found to be processed in the telencephalon. Escape reflexes control the seemingly chaotic motion of a cockroach running out from under a foot when one tries to squash it. In higher animals, examples of escape reflex include the withdrawal reflex (e.g. the withdrawal of a hand) in response to a pain stimulus. Sensory receptors in the stimulated body part send signals to the spinal cord along a sensory neuron. Within the spine, a reflex arc switches the signals straight back to the muscles of the arm (effectors) via an intermediate neuron ( interneuron) and then a motor neuron; the muscle contracts. There often is an opposite response of the opposite limb. Because this occurs automatically and indep ...
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Escape Response
Escape response, escape reaction, or escape behavior is a mechanism by which animals avoid potential predation. It consists of a rapid sequence of movements, or lack of movement, that position the animal in such a way that allows it to hide, freeze, or flee from the supposed predator. Often, an animal's escape response is representative of an instinctual defensive mechanism, though there is evidence that these escape responses may be learned or influenced by experience. The classical escape response follows this generalized, conceptual timeline: threat detection, escape initiation, escape execution, and escape termination or conclusion. Threat detection notifies an animal to a potential predator or otherwise dangerous stimulus, which provokes escape initiation, through neural reflexes or more coordinated cognitive processes. Escape execution refers to the movement or series of movements that will hide the animal from the threat or will allow for the animal to flee. Once the animal ...
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Brain
A brain is an organ that serves as the center of the nervous system in all vertebrate and most invertebrate animals. It is located in the head, usually close to the sensory organs for senses such as vision. It is the most complex organ in a vertebrate's body. In a human, the cerebral cortex contains approximately 14–16 billion neurons, and the estimated number of neurons in the cerebellum is 55–70 billion. Each neuron is connected by synapses to several thousand other neurons. These neurons typically communicate with one another by means of long fibers called axons, which carry trains of signal pulses called action potentials to distant parts of the brain or body targeting specific recipient cells. Physiologically, brains exert centralized control over a body's other organs. They act on the rest of the body both by generating patterns of muscle activity and by driving the secretion of chemicals called hormones. This centralized control allows rapid and coordinated respon ...
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Earthworm
An earthworm is a terrestrial invertebrate that belongs to the phylum Annelida. They exhibit a tube-within-a-tube body plan; they are externally segmented with corresponding internal segmentation; and they usually have setae on all segments. They occur worldwide where soil, water, and temperature allow. Earthworms are commonly found in soil, eating a wide variety of organic matter. This organic matter includes plant matter, living protozoa, rotifers, nematodes, bacteria, fungi, and other microorganisms. An earthworm's digestive system runs the length of its body. An earthworm respires (breathes) through its skin. It has a double transport system made of coelomic fluid that moves within the fluid-filled coelom and a simple, closed circulatory system. It has a central and peripheral nervous system. Its central nervous system consists of two ganglia above the mouth, one on either side, connected to a nerve running along its length to motor neurons and sensory cells in each s ...
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Amphibia
Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arboreal or freshwater aquatic ecosystems. Thus amphibians typically start out as larvae living in water, but some species have developed behavioural adaptations to bypass this. The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards but, along with mammals and birds, reptiles are amniotes and do not require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in recent decades there has been a dramatic decl ...
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Fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of living fish species are ray-finned fish, belonging to the class Actinopterygii, with around 99% of those being teleosts. The earliest organisms that can be classified as fish were soft-bodied chordates that first appeared during the Cambrian period. Although they lacked a true spine, they possessed notochords which allowed them to be more agile than their invertebrate counterparts. Fish would continue to evolve through the Paleozoic era, diversifying into a wide variety of forms. Many fish of the Paleozoic developed external armor that protected them from predators. The first fish with jaws appeared in the Silurian period, after which many (such as sharks) became formidable marine predators rather than just the prey of arthropods. Mos ...
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C-start
The Mauthner cells are a pair of big and easily identifiable neurons (one for each half of the body) located in the rhombomere 4 of the hindbrain in fish and amphibians that are responsible for a very fast escape reflex (in the majority of animals – a so-called C-start response). The cells are also notable for their unusual use of both chemical and electrical synapses. Evolutionary history Mauthner cells first appear in lampreys (being absent in hagfish and lancelets), and are present in virtually all teleost fish, as well as in amphibians (including postmetamorphic frogs and toads). Some fish, such as lumpsuckers, seem to have lost the Mauthner cells however. Role in behavior The C-start A C-start is a type of a very quick startle or escape reflex that is employed by fish and amphibians (including larval frogs and toads). There are two sequential stages in the C-start: first, the head rotates about the center of mass towards the direction of future escape, and the body ...
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Mollusca
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastropod ...
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Squid Giant Axon
The squid giant axon is the very large (up to 1.5 mm in diameter; typically around 0.5 mm) axon that controls part of the water jet propulsion system in squid. It was first described by L. W. Williams in 1909, but this discovery was forgotten until English zoologist and neurophysiologist J. Z. Young demonstrated the axon's function in the 1930s while working in the Stazione Zoologica in Naples, the Marine Biological Association in Plymouth and the Marine Biological Laboratory in Woods Hole. Squids use this system primarily for making brief but very fast movements through the water. On the underside of the squid's body, between the head and the mantle, is a siphon through which water can be rapidly expelled by the fast contractions of the body wall muscles of the animal. This contraction is initiated by action potentials in the giant axon. Action potentials travel faster in a larger axon than a smaller one, and squid have evolved the giant axon to improve the speed ...
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Crayfish
Crayfish are freshwater crustaceans belonging to the clade Astacidea, which also contains lobsters. In some locations, they are also known as crawfish, craydids, crawdaddies, crawdads, freshwater lobsters, mountain lobsters, rock lobsters, mudbugs, baybugs or yabbies. Taxonomically, they are members of the superfamilies Astacoidea and Parastacoidea. They breathe through feather-like gills. Some species are found in brooks and streams, where fresh water is running, while others thrive in swamps, ditches, and paddy fields. Most crayfish cannot tolerate polluted water, although some species, such as ''Procambarus clarkii'', are hardier. Crayfish feed on animals and plants, either living or decomposing, and detritus. The term "crayfish" is applied to saltwater species in some countries. Terminology The name "crayfish" comes from the Old French word ' (Modern French '). The word has been modified to "crayfish" by association with "fish" (folk etymology). The largely American ...
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Lateral Giant Neuron
The lateral giant interneuron (LG) is an interneuron in the abdominal nerve cord of crayfish, lobsters, shrimp of the order Decapoda and their relatives in the crustacean class Malacostraca. It is part of the system that controls a special kind of escape reflex of these organisms known as the "caridoid escape reaction." When the sensory hairs of the tail fan of crayfish are stimulated, the LG activates the motor neurons that control flexion movements of the abdomen in a way that propels the crayfish away from the source of the stimulation. The LG bypasses the main neural system that controls locomotion, thus shortening the reaction time. The lateral giant connection to motor giant fast flexor neurons was the first known example of an electrical synapse . See also *Mauthner cell *Medial giant interneuron *Squid giant axon The squid giant axon is the very large (up to 1.5 mm in diameter; typically around 0.5 mm) axon that controls part of the water jet propulsion sy ...
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Habituation
Habituation is a form of non-associative learning in which an innate (non-reinforced) response to a stimulus decreases after repeated or prolonged presentations of that stimulus. Responses that habituate include those that involve the intact organism (e.g., full-body startle response) or those that involve only components of the organism (e.g., habituation of neurotransmitter release from ''in vitro'' Aplysia sensory neurons). The broad ubiquity of habituation across all biologic phyla has resulted in it being called "the simplest, most universal form of learning...as fundamental a characteristic of life as DNA." Functionally-speaking, by diminishing the response to an inconsequential stimulus, habituation is thought to free-up cognitive resources to other stimuli that are associated with biologically important events (i.e., punishment/reward). For example, organisms may habituate to repeated sudden loud noises when they learn these have no consequences. A progressive decline of a ...
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Drosophila
''Drosophila'' () is a genus of flies, belonging to the family Drosophilidae, whose members are often called "small fruit flies" or (less frequently) pomace flies, vinegar flies, or wine flies, a reference to the characteristic of many species to linger around overripe or rotting fruit. They should not be confused with the Tephritidae, a related family, which are also called fruit flies (sometimes referred to as "true fruit flies"); tephritids feed primarily on unripe or ripe fruit, with many species being regarded as destructive agricultural pests, especially the Mediterranean fruit fly. One species of ''Drosophila'' in particular, ''D. melanogaster'', has been heavily used in research in genetics and is a common model organism in developmental biology. The terms "fruit fly" and "''Drosophila''" are often used synonymously with ''D. melanogaster'' in modern biological literature. The entire genus, however, contains more than 1,500 species and is very diverse in appearance, be ...
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