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Eubrachythoraci
Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Brachythoraci is divided into the large derived clade Eubrachythoraci and several basal groups: Buchanosteoidea, Homosteidae, and Holonematidae. (Although Holonematidae's membership in Brachythoraci is disputed.) Eubrachythoraci is then further divided into the sub-clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
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Janiosteus Timanicus
''Janiosteus'' is an extinct monospecific genus of placoderm arthrodire from the Middle Devonian: Late Givetian stage found in Timan, Russia. Phylogeny ''Janiosteus'' is a member of Panxiosteidae. In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). The phylogeny of ''Janiosteus'' within Panxiosteidae is shown in the cladogram below from the 2013 Zhu & Zhu study: However, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, instead within Coccosteomorphi and then Coccosteoidea as the sister group of Coccosteidae Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China. Phylogeny Coccosteidae belongs to the larger clade ...
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Panxiosteus Ocullus
''Panxiosteus'' is an extinct monospecific genus of placoderm arthrodire from the Middle Devonian: Givetian stage of Yunnan province, China. Phylogeny ''Panxiosteus'' is a member of Panxiosteidae. In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). The phylogeny of ''Panxiosteus'' within Panxiosteidae is shown in the cladogram below from the 2013 Zhu & Zhu study: However, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, instead within Coccosteomorphi and then Coccosteoidea as the sister group of Coccosteidae Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China. Phylogeny Coccosteidae belongs to the larger clad ...
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Coccosteomorphi
Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Coccosteomorphi is the sister taxon to Pachyosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Coccosteomorphi can be further sub-divided into Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... below: References Arthrodires {{Placoderm-stub ...
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Pachyosteomorphi
Pachyosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Pachyosteomorphi is the sister taxon to Coccosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Pachyosteomorphi can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... below: References Arthrodires {{Placoderm-stub ...
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Plourdosteus Canadensis
''Plourdosteus'' is an extinct genus of placoderm arthrodire which was relatively widespread in Euramerica during the Givetian to Frasnian ages of the Devonian. Etymology The name ''Plourdosteus'' commemorates the Plourde family at Miguasha National Park. Taxonomy ''Plourdosteus'' was previously assigned to the family Plourdosteidae within the Coccosteomorphi. However, subsequent studies found the family Plourdosteidae to be polyphyletic and should be dismissed. ''Plourdosteus'' was then proposed to be a member of Panxiosteidae. In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). The phylogeny of ''Plourdosteus'' within Panxiosteidae is shown in the cladogram below from the 2013 Zhu & Zhu study: However, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Pa ...
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Panxiosteidae
Panxiosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period. Phylogeny The family Panxiosteidae was erected by Wang in 1979. Members of the family are noted for showing morphologically intermediate traits between coccosteids and dunkleosteids. In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). The phylogeny of Panxiosteidae from the 2013 Zhu & Zhu study is shown in the cladogram below: However, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, instead within Coccosteomorphi and then Coccosteoidea as the sister group of Coccosteidae Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North Ame ...
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Brachythoraci
Brachythoraci is an extinct suborder of arthrodire placoderms, armored fish most diverse during the Devonian. Phylogeny Arthrodira is divided into three main groups: the paraphyletic Actinolepida and Phlyctaenii, and then the monophyletic Brachythoraci. Brachythoraci is then further divided into the large derived clade Eubrachythoraci and several basal groups: Buchanosteoidea, Homosteidae, and Holonematidae. (Although Holonematidae's membership in Brachythoraci is disputed.) Below is a cladogram from the 2016 Zhu ''et al.'' phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... study: References Arthrodires Prehistoric animal suborders Fish suborders {{Placoderm-stub ...
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Dunkleosteus Terrelli
''Dunkleosteus'' is an extinct genus of large armored, jawed fishes that existed during the Late Devonian period, about 382–358 million years ago. It consists of ten species, some of which are among the largest placoderms to have ever lived: ''D. terrelli'', ''D. belgicus'', ''D. denisoni'', ''D. marsaisi'', ''D. magnificus'', ''D. missouriensis'', ''D. newberryi'', ''D. amblyodoratus'', and ''D. raveri''. The largest and most well known species is ''D. terrelli'', which grew up to long and in weight. ''Dunkleosteus'' could quickly open and close its jaw, like modern-day suction feeders, and had a bite force of at the tip and at the blade edge. Numerous fossils of the various species have been found in North America, Poland, Belgium, and Morocco. Etymology ''Dunkleosteus'' was named in 1956 to honour David Dunkle (1911–1982), former curator of vertebrate paleontology at the Cleveland Museum of Natural History. The genus name ''Dunkleosteus'' combines David Dunkle's sur ...
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Dunkleosteoidea
Dunkleosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator ''Dunkleosteus terrelli'' is the best known member of this group. Phylogeny Eubrachythoraci is divided into the clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea. Dunkleosteoidea was then considered to consist of the two sister families Dunkleosteidae and Panxiosteidae. However, the 2016 Zhu ''et al.'' phylogenetic study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, leaving the status of Dunkleosteidae Dunkleosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator ''Dunkleosteus terrelli'' is the best known member of this group. Phylogeny While members of Dunkleosteidae were prev ... as a clade grouping separate from Dunkleosteoidea in doubt, as shown ...
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Arthrodire
Arthrodira (Greek for "jointed neck") is an Order (biology), order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus ...
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Aspinothoracidi
Aspinothoracidi is a clade of placoderms, extinct armored fish most diverse during the Devonian. The gigantic apex predator ''Dinichthys'', is the best-known member of this group. Many other genera, such as the infamous ''Dunkleosteus'', were previously thought to be close relatives of ''Dinichthys'' and were grouped together in the family Dinichthyidae, though more recent studies have restricted that family to only its type species. Phylogeny Eubrachythoraci is divided into the clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... below: References Arthrodires {{Placoderm-stub ...
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Protitanichthys Rockportensis
''Protitanichthys'' is an extinct genus of comparatively large coccosteid arthrodire placoderms from the Middle Devonian of the eastern United States. Fossils are found primarily in the Eifelian-epoch aged Delaware Limestone of Ohio, and the Lower Givetian-aged Rockport Quarry Limestone of Michigan Description ''Protitanichthys'' is very similar to other coccosteids, though the skull is proportionally narrower, and the orbits are comparatively smaller. As mentioned earlier, species are quite large for coccosteids, surpassed only in size by the Old World genus, '' Livosteus''. Phylogeny ''Protitanichthys'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of ''Protitanichthys'': Species ''P. fossatus'' The type species of the genus. The description of ''P. fossatus'' is based on a cranial roof, possibly long, found in the Eifelian-aged Delaware ...
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