Ctenocystoidea
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Ctenocystoidea
Ctenocystoidea is an extinct clade of echinoderms, which lived during the Cambrian and Ordovician periods. Unlike other echinoderms, ctenocystoids had bilateral symmetry, or were only very slightly asymmetrical. They are believed to be one of the earliest-diverging branches of echinoderms, with their bilateral symmetry a trait shared with other deuterostomes. Ctenocystoids were once classified in the taxon Homalozoa, also known as Carpoidea, alongside cinctans, solutes, and stylophorans. Homalozoa is now recognized as a polyphyletic group of echinoderms without radial symmetry. Ctenocystoids were geographically widespread during the Middle Cambrian, with one species surviving into the Late Ordovician. Description Like other echinoderms, ctenocystoids had a skeleton made of stereom plates. Ctenocystoids had near bilateral symmetry, with some species exhibiting slightly asymmetric plate shapes. Like cinctans and ''Ctenoimbricata'', most ctenocystoids had large marginal pla ...
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Homalozoa
Homalozoa is an obsolete extinct subphylum of Paleozoic era echinoderms, prehistoric marine invertebrates. They are also referred to as carpoids. Description The Homalozoa lacked the typical pentamer body form of other echinoderms, but all were sessile animals. Instead all Homalozoans were markedly asymmetric, and were extremely variable in forms. The body (theca) was covered with calcite plates with a number of openings. Their form is in some cases so unusual that it is unclear which openings are to be considered as mouth and anus. Many of them were stalked, similar to sea lilies (crinoids), but often their bodies were bent over, so that the mouth and anus projected forwards rather than upwards. Some forms, especially stylophorans, rested flat on the sea floor. In some forms the single ray ( brachiole or aulacophore) possessed an ambulacral groove. It has been claimed that some forms possessed gills and gill slits. Taxonomy Homalozoans were traditionally considered to be st ...
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Echinoderm
An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea lilies or "stone lilies". Adult echinoderms are found on the sea bed at every ocean depth, from the intertidal zone to the abyssal zone. The phylum contains about 7,000 living species, making it the second-largest grouping of deuterostomes, after the chordates. Echinoderms are the largest entirely marine phylum. The first definitive echinoderms appeared near the start of the Cambrian. The echinoderms are important both ecologically and geologically. Ecologically, there are few other groupings so abundant in the biotic desert of the deep sea, as well as shallower oceans. Most echinoderms are able to reproduce asexually and regenerate tissue, organs, and limbs; in some cases, they can undergo complete regeneration from a single limb. ...
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Cincta
Cincta is an extinct class of echinoderms that lived only in the Middle Cambrian epoch. Homostelea is a junior synonym. The classification of cinctans is controversial, but they are probably part of the echinoderm stem group. Cinctans were sessile, asymmetrical animals with a skeleton made of stereom plates and a racquet-shaped body composed of a theca and stele. They may have had a lifestyle similar to modern tunicates, filter-feeding by pumping water through gill slits in their pharynx. Description Cinctans were asymmetrical animals, though some species were nearly bilaterally symmetrical. Like all echinoderms, cinctans have a skeleton made of plates of stereom. The body of cinctans was divided into two sections, the main body, called the theca, and a posterior appendage called a stele. The overall shape of cinctans has been compared to a tennis racquet. The theca of cinctans was surrounded on its margins by a frame of large stereom plates called a cinctus, and the dorsal and v ...
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Ctenoimbricata
''Ctenoimbricata'' is an extinct genus of bilaterally symmetrical echinoderm, which lived during the early Middle Cambrian period of what is now Spain. It contains one species, ''Ctenoimbricata spinosa''. It may be the most basal known echinoderm. It resembles the extinct ctenocystoids and cincta Cincta is an extinct class of echinoderms that lived only in the Middle Cambrian epoch. Homostelea is a junior synonym. The classification of cinctans is controversial, but they are probably part of the echinoderm stem group. Cinctans were sessi ...ns, particularly the basal ctenocystoid ''Courtessolea''. ''Ctenoimbricata'' is interpreted as a deposit-feeding pharyngeal basket feeder. It was relatively small, with a body long. References Prehistoric echinoderm genera Cambrian echinoderms Fossil taxa described in 2012 {{cambrian-animal-stub Cambrian genus extinctions ...
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Soluta (echinoderm)
Soluta is an extinct class of echinoderms that lived from the Middle Cambrian to the Early Devonian. The class is also known by its junior synonym Homoiostelea. Soluta is one of the four "carpoid" classes, alongside Ctenocystoidea, Cincta, and Stylophora, which made up the obsolete subphylum Homalozoa. Solutes (or solutans) were asymmetric animals with a stereom skeleton and two appendages, an arm extending anteriorly and a posterior appendage called a homoiostele. Biology Most solutes were free-living, but the basal solutan ''Coleicarpus'' used its homoiostele as a holdfast, as did juvenile ''Castericystis''. Classification The phylogenetic position of Soluta is contentious. Solutans are widely agreed to be echinoderms, though the outmoded calcichordate hypothesis held that they were ancestral to both echinoderms and chordates. Within echinoderms, one hypothesis holds that stylophorans are stem-group echinoderms which branched off before echinoderms evolved radial symmetry. ...
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Drumian
The Drumian is a stage of the Miaolingian Series of the Cambrian. It succeeds the Wuliuan and precedes the Guzhangian. The base is defined as the first appearance of the trilobite ''Ptychagnostus atavus'' around million years ago. The top is defined as the first appearance of another trilobite ''Lejopyge laevigata'' around million years ago. The GSSP is defined in the ''Drumian section'' () in the Drum Mountains, Millard County, Utah, United States. The stage was also named after the Drum Mountains. The section is an outcrop of the Wheeler Formation The Wheeler Shale (named by Charles Walcott) is a Cambrian ( 507  Ma) fossil locality world-famous for prolific agnostid and ''Elrathia kingii'' trilobite remains (even though many areas are barren of fossils) and represents a Konzent ..., a succession of calcareous shales. The precise base of the Drumian is a laminated limestone above the base of the Wheeler Formation. References Cambrian geochronology Geo ...
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Cambrian Stage 4
Cambrian Stage 4 is the still unnamed fourth stage of the Cambrian and the upper stage of Cambrian Series 2. It follows Cambrian Stage 3 and lies below the Wuliuan. The lower boundary has not been formally defined by the International Commission on Stratigraphy. One proposal is the first appearance of two trilobite genera, ''Olenellus'' or '' Redlichia''. Another proposal is the first appearance of the trilobite species '' Arthricocephalus chauveaui''. Both proposals will set the lower boundary close to million years ago. The upper boundary corresponds to the beginning of the Wuliuan. Naming The International Commission on Stratigraphy The International Commission on Stratigraphy (ICS), sometimes referred to unofficially as the "International Stratigraphic Commission", is a daughter or major subcommittee grade scientific daughter organization that concerns itself with stratigr ... has not named the fourth stage of the Cambrian yet. In the widely used Siberian nomenclature stage ...
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Hemichordates
Hemichordata is a phylum which consists of triploblastic, enterocoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and include two main classes: Enteropneusta (acorn worms), and Pterobranchia. A third class, Planctosphaeroidea, is known only from the larva of a single species, ''Planctosphaera pelagica''. The class Graptolithina, formerly considered extinct, is now placed within the pterobranchs, represented by a single living genus ''Rhabdopleura''. Acorn worms are solitary worm-shaped organisms. They generally live in burrows (the earliest secreted tubes) and are deposit feeders, but some species are pharyngeal filter feeders, while the family Torquaratoridae are free living detritivores. Many are well known for their production and accumulation of various halogenated phenols and pyrroles. Pterobranchs are filter-feeders, mostly colonial, living in a collageno ...
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Blastozoa
Blastozoa is a subphylum of extinct animals belonging to Phylum Echinodermata. This subphylum is characterized by the presence of specialized respiratory structures and brachiole plates used for feeding. This subphylum ranged from the Cambrian to the Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz .... References External linksHarvard: Subphylum Blastozoa Paleozoic echinoderms Cambrian echinoderms Silurian echinoderms Ordovician echinoderms Devonian echinoderms Permian echinoderms Animal subphyla {{paleo-echinoderm-stub ...
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Periproct
The periproct is the final body segment in annelid worms. The anus is located on this segment. The term also refers to the small region surrounding the anus of the sea urchin. See also *Prostomium *Earthworm *Sea urchin Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of ... References Annelid anatomy {{animal-anatomy-stub ...
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Stereom
Stereom is a calcium carbonate material that makes up the internal skeletons found in all echinoderms, both living and fossilized forms. It is a sponge-like porous structure which, in a sea urchin may be 50% by volume living cells, and the rest being a matrix of calcite crystals. The size of openings in stereom varies in different species and in different places within the same organism. When an echinoderm becomes a fossil, microscopic examination is used to reveal the structure and such examination is often an important tool to classify the fossil as an echinoderm or related creature. Evolution Stereom was the first form of biomineralization to evolve in deuterostomes, predating the evolution of spicules in tunicates and bone in vertebrates. Stereom likely evolved before other distinctive traits of echinoderms, such as radial symmetry, as it is present in basal echinoderms with bilaterally symmetric or asymmetric body plans. Stereom may have evolved alongside a transition in ocea ...
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