Chasmataspidid
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Chasmataspidid
Chasmataspidids, sometime referred to as chasmataspids, are a group of extinct chelicerate arthropods that form the order Chasmataspidida. Chasmataspidids are probably related to horseshoe crabs (Xiphosura) and/or sea scorpions (Eurypterida), with more recent studies suggest that they form a clade (Dekatriata) with Eurypterida and Arachnida. Chasmataspidids are known sporadically in the fossil record through to the mid-Devonian, with possible evidence suggesting that they were also present during the late Cambrian. Chasmataspidids are most easily recognised by having an opisthosoma divided into a wide forepart (preabdomen) and a narrow hindpart (postabdomen) each comprising 4 and 9 segments respectively. There is some debate about whether they form a natural (i.e. monophyletic) group. Distribution Chasmataspidids survived at least since Ordovician to mid-Devonian in age. As of 2019, most chasmataspidids (with a total of 9 species) are known from the Devonian strata, while the p ...
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Dvulikiaspis
''Dvulikiaspis'' is a genus of chasmataspidid, a group of extinct aquatic arthropods. Fossils of the single and type species, ''D. menneri'', have been discovered in deposits of the Early Devonian period (Lochkovian epoch) in the Krasnoyarsk Krai, Siberia, Russia. The name of the genus is composed by the Russian word двуликий (''dvulikij''), meaning "two-faced", and the Ancient Greek word ἀσπίς ('' aspis''), meaning "shield". The species name honors the discoverer of the holotype of ''Dvulikiaspis'', Vladimir Vasilyevich Menner. Its prosoma (head) was subquadrate (almost square) to parabolic (nearly U-shaped), with (bean-shaped) to subovate (nearly oval) eyes and surrounded by a marginal rim. The abdomen was composed by a fused buckler and a postabdomen that occupied most of the body length, while the telson (the posteriormost division of the body) was small and semicircular in shape. The appendages (limbs) were uniform. The sixth and last pair of them had a pad ...
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Chelicerates
The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. The Chelicerata originated as marine animals in the Middle Cambrian period; the first confirmed chelicerate fossils, belonging to ''Sanctacaris'', date from 508 million years ago. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered ...
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Chelicerate
The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. The Chelicerata originated as marine animals in the Middle Cambrian period; the first confirmed chelicerate fossils, belonging to '' Sanctacaris'', date from 508 million years ago. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered ...
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Chasmataspis
''Chasmataspis'' is a genus of chasmataspidid, a group of extinct aquatic chelicerate arthropods. It was found in the Early Ordovician deposits of Tennessee, USA. Morphology Just like other chasmataspidids, the body of ''Chasmataspis'' compose of a prosoma and a 13-segmented opisthosoma, with the latter subdivided into a 4-segmented preabdomen and a 9-segmented postabdomen. Dorsal to the carapace were pairs of ridges, lateral eyes and median ocelli. ''Chasmataspis'' characterized by a semicircular carapace (prosomal dorsal shield) with pointed genal spines and preabdomen with fused body segments. Rows of tubercles and spines run through the axial and lateral regions of opisthosoma. The body terminated with a spine-like telson. There is no body fossils of ''Chasmataspis'' with unambiguous appendages being discovered. The only evidence were 2 specimens of disarticulated, claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mam ...
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Chasmataspididae
''Chasmataspis'' is a genus of chasmataspidid, a group of extinct aquatic chelicerate arthropods. It was found in the Early Ordovician deposits of Tennessee, USA. Morphology Just like other chasmataspidids, the body of ''Chasmataspis'' compose of a prosoma and a 13-segmented opisthosoma, with the latter subdivided into a 4-segmented preabdomen and a 9-segmented postabdomen. Dorsal to the carapace were pairs of ridges, lateral eyes and median ocelli. ''Chasmataspis'' characterized by a semicircular carapace (prosomal dorsal shield) with pointed genal spines and preabdomen with fused body segments. Rows of tubercles and spines run through the axial and lateral regions of opisthosoma. The body terminated with a spine-like telson. There is no body fossils of ''Chasmataspis'' with unambiguous appendages being discovered. The only evidence were 2 specimens of disarticulated, claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mam ...
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Kiaeria Limuloides
''Kiaeria'' is a genus of chasmataspidid, a group of extinct aquatic arthropods. It was originally classified as a xiphosuran of the monotypic family Kiaeriidae. However, in 2019, the British geologist and paleobiologist James C. Lamsdell assigned the genus to the order Chasmataspidida, possibly being a member of the family Chasmataspididae. Only one species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ... has been assigned to this genus, ''K. limuloides''. References Prehistoric chelicerates Chasmataspidida {{Paleo-arthropod-stub ...
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Hoplitaspis
''Hoplitaspis'' is a genus of chasmataspidid, an extinct group of aquatic arthropods. Fossils of ''Hoplitaspis'' have been discovered in Late Ordovician deposits of the Lagerstätte of the Big Hill Formation exposed at Stonington Peninsular in Michigan's Upper Peninsula, United States. Its generic name is derived from the hoplites (Ancient Greek citizen-soldiers) and the Ancient Greek word άσπίς ('' aspis'', meaning "shield"). The specific name ''hiawathai'' honors Hiawatha Hiawatha ( , also : ), also known as Ayenwathaaa or Aiionwatha, was a precolonial Native American leader and co-founder of the Iroquois Confederacy. He was a leader of the Onondaga people, the Mohawk people, or both. According to some account ..., a Native American leader and co-founder of the Iroquois Confederacy. References Diploaspididae Late Ordovician first appearances Ordovician chasmataspidids Chasmataspidids of North America Fossils of the United States Fossil taxa described ...
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Dekatriata
Dekatriata is a clade of planatergan chelicerates including the groups Arachnida, Chasmataspidida, Eurypterid Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is l ...a and additionally two stem-genera '' Winneshiekia'' and '' Houia''. Dekatriata is defined by an opisthosoma with 13 segments as groundplan (the number proposed to be secondarily reduced in most arachnid orders) and fused, plate-like appendages on the first opisthosomal segment (somite VII). References Middle Ordovician first appearances {{Chelicerata-stub ...
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Eurypterid
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 Myr, million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic Chelicerata, chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Tri ...
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Eurypterida
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event (or sometim ...
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Xiphosura
Xiphosura () is an order of arthropods related to arachnids. They are more commonly known as horseshoe crabs (a name applied more specifically to the only extant family, Horseshoe crab, Limulidae). They first appeared in the Hirnantian (Late Ordovician). Currently, there are only four living species. Xiphosura contains one suborder, Xiphosurida, and several stem-genera. The group has hardly changed in appearance in hundreds of millions of years; the modern horseshoe crabs look almost identical to prehistoric genera and are considered to be living fossils. The most notable difference between ancient and modern forms is that the abdominal segments in present species are fused into a single unit in adults. Xiphosura were historically placed in the class Merostomata, although this term was intended to encompass also the Eurypterida, eurypterids, whence it denoted what is now known to be an unnatural (paraphyletic) group (although this is a grouping recovered in some recent cladistic ...
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Prosoma
The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cephalothorax'' and ''abdomen'' in some groups.) The word ''cephalothorax'' is derived from the Greek words for head (, ') and thorax (, '). This fusion of the head and thorax is seen in chelicerates and crustaceans; in other groups, such as the Hexapoda (including insects), the head remains free of the thorax. In horseshoe crabs and many crustaceans, a hard shell called the carapace covers the cephalothorax. Arachnid anatomy Fovea The fovea is the centre of the cephalothorax and is located behind the head (only in spiders).Dalton, Steve (2008). ''Spiders; The Ultimate Predators''. A & C Black, London. P.p. 19. . It is often important in identification. It can be transverse or procurved Smith, A. M. (1990c). Baboon spiders: Tarantulas of Afr ...
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