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Chimpanzee–human Last Common Ancestor
The chimpanzee–human last common ancestor (CHLCA) is the last common ancestor shared by the extant ''Homo'' (human) and '' Pan'' (chimpanzee and bonobo) genera of Hominini. Due to complex hybrid speciation, it is not currently possible to give a precise estimate on the age of this ancestral population. While "original divergence" between populations may have occurred as early as 13 million years ago (Miocene), hybridization may have been ongoing until as recently as 4 million years ago (Pliocene). In human genetic studies, the CHLCA is useful as an anchor point for calculating single-nucleotide polymorphism (SNP) rates in human populations where chimpanzees are used as an outgroup, that is, as the extant species most genetically similar to ''Homo sapiens''. Taxonomy The taxon tribe ''Hominini'' was proposed to separate humans (genus ''Homo'') from chimpanzees ('' Pan'') and gorillas (genus ''Gorilla'') on the notion that the least similar species should be separat ...
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Hominini Lineage
The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The term was originally introduced by Camille Arambourg (1948). Arambourg combined the categories of ''Hominina'' and ''Simiina'' due to Gray (1825) into his new subtribe. Traditionally, chimpanzees, gorillas and orangutans were grouped together as pongids. Since Gray's classification, evidence has accumulated from genetic phylogeny confirming that humans, chimpanzees, and gorillas are more closely related to each other than to the orangutan. The former pongids were reassigned to the subfamily Hominidae ("great apes"), which already included humans, but the details of this reassignment remain contested; within Hominini, not every source excludes gorillas, and not every source includes chimpanzees. Humans are the only extant species in the A ...
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Panina
The genus ''Pan'' consists of two extant species: the chimpanzee and the bonobo. Taxonomically, these two ape species are collectively termed panins; however, both species are more commonly referred to collectively using the generalized term chimpanzees, or chimps. Together with humans, gorillas, and orangutans they are part of the family Hominidae (the great apes, or ''hominids''). Native to sub-Saharan Africa, chimpanzees and bonobos are currently both found in the Congo jungle, while only the chimpanzee is also found further north in West Africa. Both species are listed as endangered on the IUCN Red List of Threatened Species, and in 2017 the Convention on Migratory Species selected the chimpanzee for special protection. Chimpanzee and bonobo: comparison The chimpanzee (''P. troglodytes'') who lives north of the Congo River, and the bonobo (''P. paniscus'') who lives south of it, were once considered to be the same species, but since 1928 they have been recognized as di ...
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Nature (journal)
''Nature'' is a British weekly scientific journal founded and based in London, England. As a multidisciplinary publication, ''Nature'' features peer-reviewed research from a variety of academic disciplines, mainly in science and technology. It has core editorial offices across the United States, continental Europe, and Asia under the international scientific publishing company Springer Nature. ''Nature'' was one of the world's most cited scientific journals by the Science Edition of the 2019 ''Journal Citation Reports'' (with an ascribed impact factor of 42.778), making it one of the world's most-read and most prestigious academic journals. , it claimed an online readership of about three million unique readers per month. Founded in autumn 1869, ''Nature'' was first circulated by Norman Lockyer and Alexander Macmillan as a public forum for scientific innovations. The mid-20th century facilitated an editorial expansion for the journal; ''Nature'' redoubled its efforts in exp ...
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Ardipithecus
''Ardipithecus'' is a genus of an extinct hominine that lived during the Late Miocene and Early Pliocene epochs in the Afar Depression, Ethiopia. Originally described as one of the earliest ancestors of humans after they diverged from the chimpanzees, the relation of this genus to human ancestors and whether it is a hominin is now a matter of debate. Two fossil species are described in the literature: '' A. ramidus'', which lived about 4.4 million years ago during the early Pliocene, and '' A. kadabba'', dated to approximately 5.6 million years ago (late Miocene). Initial behavioral analysis indicated that ''Ardipithecus'' could be very similar to chimpanzees, however more recent analysis based on canine size and lack of canine sexual dimorphism indicates that ''Ardipithecus'' was characterised by reduced aggression, and that they more closely resemble bonobos. Some analyses describe ''Australopithecus'' as being sister to ''Ardipithecus ramidus'' specifically. This mea ...
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Hominine
Homininae (), also called "African hominids" or "African apes", is a subfamily of Hominidae. It includes two tribes, with their extant as well as extinct species: 1) the tribe Hominini (with the genus ''Homo'' including modern humans and numerous extinct species; the subtribe Hominina, comprising at least two extinct genera; and the subtribe Panina, represented only by the genus ''Pan'', which includes chimpanzees and bonobos)―and 2) the tribe Gorillini (gorillas). Alternatively, the genus '' Pan'' is sometimes considered to belong to its own third tribe, Panini. Homininae comprises all hominids that arose after orangutans (subfamily Ponginae) split from the line of great apes. The Homininae cladogram has three main branches, which lead to gorillas (through the tribe Gorillini), and to humans and chimpanzees via the tribe Hominini and subtribes Hominina and Panina (see the evolutionary tree below). There are two living species of Panina (chimpanzees and bonobos) and two living ...
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Sahelanthropus Tchadensis
''Sahelanthropus tchadensis'' is an extinct species of the Homininae (African apes) dated to about , during the Miocene epoch. The species, and its genus ''Sahelanthropus'', was announced in 2002, based mainly on a partial cranium, nicknamed ''Toumaï'', discovered in northern Chad. ''Sahelanthropus tchadensis'' lived close to the time of the chimpanzee–human divergence, possibly related to ''Orrorin'', a species of Homininae that lived about one million years later. It may have been ancestral to both humans and chimpanzees (which would place it in the tribe Hominini), or alternatively an early member of the tribe Gorillini. In 2020, the femur was analyzed, and it was found that ''Sahelanthropus'' was not bipedal, casting some doubt on its position as a human ancestor, but this was refuted in 2022. Taxonomy Discovery Funded by the Mission Paléoanthropologique Franco–Tchadienne, Ahounta Djimdoumalbaye, Fanoné Gongdibé, Mahamat Adoum, and Alain Beauvilain identified the f ...
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Southeast Europe
Southeast Europe or Southeastern Europe (SEE) is a geographical subregion of Europe, consisting primarily of the Balkans. Sovereign states and territories that are included in the region are Albania, Bosnia and Herzegovina, Bulgaria, Croatia (alternatively placed in Central Europe), Cyprus (alternatively placed in West Asia), Greece (alternatively placed in Southern Europe), Kosovo, Montenegro, North Macedonia, Romania, Serbia, and Turkey (alternatively placed in Southern Europe or West Asia). Sometimes, Moldova (alternatively placed in Eastern Europe) and Slovenia (alternatively placed in Central Europe) are also included. The largest city of the region is Istanbul, followed by Bucharest, Sofia, Belgrade, and Athens. There are overlapping and conflicting definitions of the region, due to political, economic, historical, cultural, and geographical considerations. Definition The first known use of the term "Southeast Europe" was by Austrian researcher Johann Georg von Hahn (1811â ...
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Graecopithecus
''Graecopithecus'' is an extinct species of hominid that lived in southeast Europe during the late Miocene around 7.2 million years ago. Originally identified by a single lower jaw bone bearing a molar tooth found in Pyrgos Vasilissis, Athens, Greece, in 1944, other tooth specimens were discovered from Azmaka quarry in Bulgaria in 2012. With only little and badly preserved materials to reveal its nature, it is considered as "the most poorly known European Miocene hominoids." The creature was popularly nicknamed 'El Graeco' (word play on the Spanish painter El Greco) by scientists. In 2017, an international team of palaeontologists led by Madelaine Böhme of the Eberhard-Karls-University Tübingen, Germany, published a detailed analysis of the teeth and age of the specimens, and came to the conclusion that it could be the oldest hominin, meaning that it could be the oldest direct ancestors of humans after splitting from that of the chimpanzees. Their simultaneous study also clai ...
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Hominidae
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the eastern and western gorilla); '' Pan'' (the chimpanzee and the bonobo); and ''Homo'', of which only modern humans remain. Several revisions in classifying the great apes have caused the use of the term ''"hominid"'' to vary over time. The original meaning of "hominid" referred only to humans (''Homo'') and their closest extinct relatives. However, by the 1990s humans, apes, and their ancestors were considered to be "hominids". The earlier restrictive meaning has now been largely assumed by the term ''"hominin"'', which comprises all members of the human clade after the split from the chimpanzees (''Pan''). The current meaning of "hominid" includes all the great apes including humans. Usage still varies, however, and some scientists and la ...
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Australopithecus
''Australopithecus'' (, ; ) is a genus of early hominins that existed in Africa during the Late Pliocene and Early Pleistocene. The genus ''Homo'' (which includes modern humans) emerged within ''Australopithecus'', as sister to e.g. ''Australopithecus sediba''. Also the genera ''Paranthropus'' and ''Kenyanthropus'' emerged within ''Australopithecus''. ''Australopithecus'' is a member of the subtribe Australopithecina, which sometimes also includes ''Ardipithecus'', though the term "australopithecine" is sometimes used to refer only to members of ''Australopithecus''. Species include ''Australopithecus garhi, A. garhi'', ''Australopithecus africanus, A. africanus'', ''Australopithecus sediba, A. sediba'', ''Australopithecus afarensis, A. afarensis, Australopithecus anamensis, A. anamensis, Australopithecus bahrelghazali, A. bahrelghazali'' and ''Australopithecus deyiremeda, A. deyiremeda''. Debate exists as to whether some ''Australopithecus'' species should be reclassified into ne ...
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Pan Troglodytes
The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of Hominidae, great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative the bonobo was more commonly known as the pygmy chimpanzee, this species was often called the common chimpanzee or the robust chimpanzee. The chimpanzee and the bonobo are the only species in the genus Pan (genus), ''Pan''. Evidence from fossils and DNA sequencing shows that ''Pan'' is a sister taxon to the Human evolution, human lineage and is humans' closest living relative. The chimpanzee is covered in coarse black hair, but has a bare face, fingers, toes, palms of the hands, and soles of the feet. It is larger and more Robustness (morphology), robust than the bonobo, weighing for males and for females and standing . The chimpanzee lives in groups that range in size from 15 to 150 members, although individuals travel and forag ...
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Richard Wrangham
Richard Walter Wrangham (born 1948) is an English anthropologist and primatologist; he is Professor of Biological Anthropology at Harvard University. His research and writing have involved ape behavior, human evolution, violence, and cooking. Biography Wrangham was born in Leeds, Yorkshire. Following his years on the faculty of the University of Michigan, he became the Ruth Moore Professor of Biological Anthropology at Harvard University and his research group is now part of the newly established Department of Human Evolutionary Biology. He is a MacArthur fellow. He is co-director of the Kibale Chimpanzee Project, the long-term study of the Kanyawara chimpanzees in Kibale National Park, Uganda. His research culminates in the study of human evolution in which he draws conclusions based on the behavioural ecology of apes. As a graduate student, Wrangham studied under Robert Hinde and Jane Goodall. Wrangham is known predominantly for his work in the ecology of primate social s ...
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