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Carnosaurs
Carnosauria is an extinct large group of predatory dinosaurs that lived during the Jurassic and Cretaceous periods. Starting from the 1990s, scientists have discovered some very large carnosaurs in the carcharodontosaurid family, such as ''Giganotosaurus'', ''Mapusaurus'', ''Carcharodontosaurus'' and ''Tyrannotitan'' which are among the largest known predatory dinosaurs. While it originally contained a wide assortment of giant theropods that were not closely related, the group has since been defined to encompass only the allosaurs and their closest kin. However, with the description and publication in 2019 of ''Asfaltovenator vialidadi'', a basal allosauroid curiously displaying both primitive and derived features seen in Tetanurae, the new phylogenetic analysis has found Megalosauroidea to be a basal grade of carnosaurs in respect to Allosauroidea; thus significantly expanding Carnosauria's inclusiveness towards its original context. Distinctive characteristics of carnosaurs ...
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Tetanurae
Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans (including birds). Tetanurans are defined as all theropods more closely related to modern birds than to ''Ceratosaurus'' and contain the majority of predatory dinosaur diversity. Tetanurae likely diverged from its sister group, Ceratosauria, during the late Triassic. Tetanurae first appeared in the fossil record by the Early Jurassic about 190 mya and by the Middle Jurassic had become globally distributed. The group was named by Jacques Gauthier in 1986 and originally had two main subgroups: Carnosauria and Coelurosauria, the clade containing birds and related dinosaurs such as compsognathids, tyrannosaurids, ornithomimosaurs, and maniraptorans. The original Carnosauria was a polyphyletic group including any large carnivorous theropod. Many of Gauthier's carnosaurs, such ...
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Theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic period 231.4 million years ago ( Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of diets was historically considered a characteri ...
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Megalosauridae
Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relatively primitive group of basal tetanurans containing two main subfamilies, Megalosaurinae and Afrovenatorinae, along with the basal genus ''Eustreptospondylus'', an unresolved taxon which differs from both subfamilies. The defining megalosaurid, ''Megalosaurus bucklandii'', was first named and described in 1824 by William Buckland after multiple finds in Stonesfield, Oxfordshire, UK. ''Megalosaurus'' was the first formally described dinosaur and was the basis for the establishment of the clade Dinosauria. It is also one of the largest known Middle Jurassic carnivorous dinosaurs, with the best-preserved femur at 805 mm and a proposed body mass of around 943 kg. Megalosauridae has mainly been recognized as a European group of dinosaurs, ...
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Spinosauridae
The Spinosauridae (or spinosaurids) are a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. They came into prominence during the Cretaceous period. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America and Asia. Their remains have generally been attributed to the Early to Mid Cretaceous. Spinosaurids were large bipedal carnivores. Their crocodilian-like skulls were long, low and narrow, bearing conical teeth with reduced or absent serrations. The tips of their upper and lower jaws fanned out into a spoon-shaped structure similar to a rosette, behind which there was a notch in the upper jaw that the expanded tip of the lower jaw fit into. The nostrils of spinosaurids were retracted to a position further back on the head than in most other theropods, and they had bony crests on their heads along the midline of their skulls. Their robust shoulders wielded stocky forelimbs, with three-fingered hands that b ...
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Megalosauroidea
Megalosauroidea (meaning 'great/big lizard forms') is a superfamily (or clade) of tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as ''Megalosaurus bucklandii'' and all taxa sharing a more recent common ancestor with it than with ''Allosaurus fragilis'' or ''Passer domesticus''. Members of the group include ''Spinosaurus'', ''Megalosaurus'', and ''Torvosaurus''. They are possibly paraphyletic in nature. Classification The name Spinosauroidea is sometimes used in place of Megalosauroidea. The superfamily Spinosauroidea was named in 1915 by Ernst Stromer. It is a synonym of Megalosauroidea in almost all modern phylogenetic analyses, and it is therefore redundant. Spinosauroidea was defined as a clade in 1998 by Paul Sereno as the node clade containing the common ancestor of ''Spinosaurus'' and ''Torvosaurus'' and all its descendants. Thomas Holtz in 2004 defined a branch clade with the same name containing all spec ...
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Allosauroidea
Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four family (biology), families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroidea, were among the apex predators that were active during the Middle Jurassic to Late Cretaceous periods. The most famous and best understood allosauroid is the North American genus ''Allosaurus''. The oldest-known allosauroid, ''Shidaisaurus jinae'', appeared in the early Middle Jurassic about 174 million years ago (Earliest Aalenian stage) of China. The last known definitive surviving members of the group died out around 89 million years ago in Asia (''Shaochilong'') and South America (''Mapusaurus''), though the megaraptorans, which survived until the end of Maastrichtian, may belong to the group as well. A frontal assigned to an allosauroid found to be most closely related to ''Sinraptor'' has also been found in the Coniacian (89-86.3 ...
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Piatnitzkysauridae
Piatnitzkysauridae is an extinct family of megalosauroid or basal Allosauroid dinosaurs. It only consists of three known dinosaur genera: '' Condorraptor'', ''Marshosaurus'', and ''Piatnitzkysaurus.'' The most complete and well known member of this family is ''Piatnitzkysaurus'', which also gives the family its name. Description So far all known piatnitzkysaurids have only been found in Jurassic deposits of the western hemisphere/New world. ''Piatnitzkysaurus'' and ''Condorraptor'' hail from the Cañadón Asfalto Formation of Argentina, which has been dated from the Toarcian epoch of the Early Jurassic to the Bathonian epoch of the Middle Jurassic (approximately 179 to 168 million years ago). ''Marshosaurus'' was found in the Morrison Formation of the United States, which was dated to the Kimmeridgian epoch of the Late Jurassic. Piatnitzkysaurids were among the first large theropods present in South America, and are evidence for a radiation of basal tetanurans in the middle Jur ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, are among ...
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Carcharodontosaurus
''Carcharodontosaurus'' (; ) is a genus of large carcharodontosaurid theropod dinosaur that existed during the Cenomanian age of the Late Cretaceous in Northern Africa. The genus ''Carcharodontosaurus'' is named after the shark genus ''Carcharodon'', itself composed of the Greek (, meaning "jagged" or "sharp") and (, "teeth"), and the suffix ' ("lizard"). It is currently known to have two species: ''C. saharicus'' and ''C. iguidensis''. History of discovery In 1924, two teeth were found in the Continental intercalaire of Algeria, showing what were at the time unique characteristics. These teeth were described by Depéret and Savornin (1925) as representing a new taxon, which they named ''Megalosaurus saharicus'' and later categorized in the subgenus ''Dryptosaurus''. Some years later, paleontologist Ernst Stromer described the remains of a partial skull and skeleton from Cenomanian aged rocks in the Bahariya Formation of Egypt (Stromer, 1931); originally excavated in 1914, ...
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Giganotosaurus
''Giganotosaurus'' ( ) is a genus of theropod dinosaur that lived in what is now Argentina, during the early Cenomanian age of the Late Cretaceous period, approximately 99.6 to 95 million years ago. The holotype specimen was discovered in the Candeleros Formation of Patagonia in 1993 and is almost 70% complete. The animal was named ''Giganotosaurus carolinii'' in 1995; the genus name translates to "giant southern lizard", and the specific name honors the discoverer, Rubén D. Carolini. A dentary bone, a tooth, and some tracks, discovered before the holotype, were later assigned to this animal. The genus attracted much interest and became part of a scientific debate about the maximum sizes of theropod dinosaurs. ''Giganotosaurus'' was one of the largest known terrestrial carnivores, but the exact size has been hard to determine due to the incompleteness of the remains found so far. Estimates for the most complete specimen range from a length of , a skull in length, and a we ...
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Tyrannotitan
''Tyrannotitan'' (; ) is a genus of huge bipedal carnivorous dinosaur of the carcharodontosaurid family from the Aptian stage of the early Cretaceous period, discovered in Argentina. It is closely related to other giant predators like ''Carcharodontosaurus'' and especially ''Giganotosaurus'' as well as ''Mapusaurus''. Discovery and species ''Tyrannotitan chubutensis'' was described by Fernando E. Novas, Silvina de Valais, Pat Vickers-Rich, and Tom Rich in 2005. The fossils were found at La Juanita Farm, northeast of Paso de Indios, Chubut Province, Argentina. They are believed to have been from the Cerro Castaño Member, Cerro Barcino Formation (Aptian stage). The holotype material was designated MPEF-PV 1156 and included partial dentaries, teeth, back vertebrae 3–8 and 11–14, proximal tail vertebrae, ribs and chevrons, a fragmentary scapulocoracoid, humerus, ulna, partial ilium, a nearly complete femur, fibula, and left metatarsal 2. Additional material (designated M ...
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Asfaltovenator
''Asfaltovenator'' (meaning "Cañadón Asfalto Formation hunter" after the fossil formation in which its fossils were found) is a genus of possibly allosauroid dinosaur from the Lower Jurassic Cañadón Asfalto Formation from Chubut Province, Argentina. The type and only species is ''Asfaltovenator vialidadi''. Discovery In 2002, technician Leandro Canesa, roughly a mile north of Cerro Condor, discovered a theropod skeleton. Excavations started in 2005. In 2007, the fossil was in its entirety removed as an enormous stone block, covered by plaster of Paris. It was then prepared by Mariano Caffa, a process that took five years due to the extreme hardness of the stone matrix. Between 2013 and 2015, it was compared with the specimens of related theropods, researchers personally investigating their exemplars in collections all over the world. In 2019, the type species ''Asfaltovenator vialidadi'' was named and described by Oliver Walter Mischa Rauhut and Diego Pol. The generic name ...
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