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Bipinnaria
A bipinnaria is the first stage in the larval development of most starfish, and is usually followed by a brachiolaria stage. Movement and feeding is accomplished by the bands of cilia. Starfish that brood their young generally lack a bipinnaria stage, with the eggs developing directly into miniature adults The bipinnaria is free-living, swimming as part of the zooplankton. When it initially forms, the entire body is covered by cilia, but as it grows, these become confined to a narrow band forming a number of loops over the body surface. A pair of short, stubby arms soon develop on the body, with the ciliated bands extending into them. In addition to propelling the larva through the water, the cilia also catch suspended food particles, and deliver them to the mouth (more correctly called a ''stomodeum''). Eventually, three additional arms develop at the front end of the larva; at this point it becomes a brachiolaria. In some species, including the common starfish ''Asterias'', ...
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Starfish
Starfish or sea stars are star-shaped echinoderms belonging to the class Asteroidea (). Common usage frequently finds these names being also applied to ophiuroids, which are correctly referred to as brittle stars or basket stars. Starfish are also known as asteroids due to being in the class Asteroidea. About 1,900 species of starfish live on the seabed in all the world's oceans, from warm, tropical zones to frigid, polar regions. They are found from the intertidal zone down to abyssal depths, at below the surface. Starfish are marine invertebrates. They typically have a central disc and usually five arms, though some species have a larger number of arms. The aboral or upper surface may be smooth, granular or spiny, and is covered with overlapping plates. Many species are brightly coloured in various shades of red or orange, while others are blue, grey or brown. Starfish have tube feet operated by a hydraulic system and a mouth at the centre of the oral or lower surface. ...
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Brachiolaria
A brachiolaria is the second stage of larval development in many starfishes. It follows the bipinnaria. Brachiolaria have bilateral symmetry, unlike the adult starfish, which have a pentaradial symmetry. Starfish of the order Paxillosida (''Astropecten'' and '' Asterina'') have no brachiolaria stage, with the bipinnaria developing directly into an adult. The brachiolaria develops from the bipinnaria larva when the latter grows three short arms at the underside of its anterior end. These arms each bear sticky cells at the tip, and they surround an adhesive sucker. The larva soon sinks to the bottom, attaching itself to the substrate, firstly with the tips of the arms, and then with the sucker. Once attached, it begins to metamorphose into the adult form. The adult starfish develops only from the hind-part of the larva, away from the sucker. It is from this part that the arms of the adult grow, with the larval arms eventually degenerating and disappearing. The digestive system of the ...
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Larva
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The larva's appearance is generally very different from the adult form (''e.g.'' caterpillars and butterflies) including different unique structures and organs that do not occur in the adult form. Their diet may also be considerably different. Larvae are frequently adapted to different environments than adults. For example, some larvae such as tadpoles live almost exclusively in aquatic environments, but can live outside water as adult frogs. By living in a distinct environment, larvae may be given shelter from predators and reduce competition for resources with the adult population. Animals in the larval stage will consume food to fuel their transition into the adult form. In some organisms like polychaetes and barnacles, adults are immobil ...
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Larva Bipinnaria
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The larva's appearance is generally very different from the adult form (''e.g.'' caterpillars and butterflies) including different unique structures and organs that do not occur in the adult form. Their diet may also be considerably different. Larvae are frequently adapted to different environments than adults. For example, some larvae such as tadpoles live almost exclusively in aquatic environments, but can live outside water as adult frogs. By living in a distinct environment, larvae may be given shelter from predators and reduce competition for resources with the adult population. Animals in the larval stage will consume food to fuel their transition into the adult form. In some organisms like polychaetes and barnacles, adults are immo ...
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Tornaria
A tornaria is the planktonic larva of some species of Hemichordata such as the acorn worms. It is very similar in appearance to the bipinnaria larvae of starfishes, with convoluted bands of cilia The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projecti ... running around the body. It is an oval shaped, transparent larva. The diameter of the body is about 3 mm. It has an apical plate, which is a thickened region provided by a tuft of cilia and a pair of eye spots. The larva has a complete alimentary canal. The ciliary band stretches throughout the anterior and posterior region, and also the postoral region. References Larvae Hemichordates {{biology-stub ...
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Echinoderm Biology
An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea lilies or "stone lilies". Adult echinoderms are found on the sea bed at every ocean depth, from the intertidal zone to the abyssal zone. The phylum contains about 7,000 living species, making it the second-largest grouping of deuterostomes, after the chordates. Echinoderms are the largest entirely marine phylum. The first definitive echinoderms appeared near the start of the Cambrian. The echinoderms are important both ecologically and geologically. Ecologically, there are few other groupings so abundant in the biotic desert of the deep sea, as well as shallower oceans. Most echinoderms are able to reproduce asexually and regenerate tissue, organs, and limbs; in some cases, they can undergo complete regeneration from a single limb. Geologi ...
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Cilia
The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projection that extends from the surface of the much larger cell body. Eukaryotic flagella found on sperm cells and many protozoans have a similar structure to motile cilia that enables swimming through liquids; they are longer than cilia and have a different undulating motion. There are two major classes of cilia: ''motile'' and ''non-motile'' cilia, each with a subtype, giving four types in all. A cell will typically have one primary cilium or many motile cilia. The structure of the cilium core called the axoneme determines the cilium class. Most motile cilia have a central pair of single microtubules surrounded by nine pairs of double microtubules called a 9+2 axoneme. Most non-motile cilia have a 9+0 axoneme that lacks the central pair of mi ...
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Zooplankton
Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by currents in the ocean, or by currents in seas, lakes or rivers. Zooplankton can be contrasted with phytoplankton, which are the plant component of the plankton community ("phyto" comes from the Greek word for ''plant''). Zooplankton are heterotrophic (other-feeding), whereas phytoplankton are autotrophic (self-feeding). This means zooplankton cannot manufacture their own food but must eat other plants or animals instead — in particular they eat phytoplankton. Zooplankton are generally larger than phytoplankton, most are microscopic, but some (such as jellyfish) are macroscopic and can be seen with the naked eye. Many protozoans (single-celled protists that prey on other microscopic life) are zooplankton, including zooflagellates, fo ...
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Asterias
''Asterias'' is a genus of the Asteriidae family of sea stars. It includes several of the best-known species of sea stars, including the (Atlantic) common starfish, ''Asterias rubens'', and the northern Pacific seastar, ''Asterias amurensis''. The genus contains a total of eight species in all. All species have five arms and are native to shallow oceanic areas (the littoral zone) of cold to temperate parts of the Holarctic. These starfish have planktonic larvae. ''Asterias amurensis'' is an invasive species in Australia and can in some years become a pest in the Japanese mariculture industry. History The genus ''Asterias'' was first described by Carl Linnaeus in the 10th edition of Systema Naturae in 1758 when he published ''A. rubens''. It was for a time the only species, but by the early 1800s a few dozen taxa had been described in this genus. In 1825 Thomas Say listed six species native to the coasts of the United States (which at the time consisted of the east coast from M ...
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Hemichordates
Hemichordata is a phylum which consists of triploblastic, enterocoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and include two main classes: Enteropneusta (acorn worms), and Pterobranchia. A third class, Planctosphaeroidea, is known only from the larva of a single species, ''Planctosphaera pelagica''. The class Graptolithina, formerly considered extinct, is now placed within the pterobranchs, represented by a single living genus ''Rhabdopleura''. Acorn worms are solitary worm-shaped organisms. They generally live in burrows (the earliest secreted tubes) and are deposit feeders, but some species are pharyngeal filter feeders, while the family Torquaratoridae are free living detritivores. Many are well known for their production and accumulation of various halogenated phenols and pyrroles. Pterobranchs are filter-feeders, mostly colonial, living in a collageno ...
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Ambulacraria
Ambulacraria , or Coelomopora , is a clade of invertebrate phyla that includes echinoderms and hemichordates; a member of this group is called an ambulacrarian. Phylogenetic analysis suggests the echinoderms and hemichordates separated around 533 million years ago. The Ambulacraria are part of the deuterostomes, a larger clade that also includes the Chordata, Vetulicolia. The two living clades with representative organisms are: * Echinodermata (sea stars, sea urchins, brittle stars, sea cucumbers, feather stars, sea lilies, etc.) * Hemichordata (acorn worms, Pterobranchia, and possibly graptolites) (These together sometimes are called the ''lower deuterostomes''.) Whether the Xenacoelomorpha clade is the sister group to the Ambulacraria remains a contentious issue, with some authors arguing that the former should be placed more basally among metazoans, and other authors asserting that the best choices of phylogenetic methods support the position of Xenacoelomorpha as the sister ...
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