Bilaterian
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Bilaterian
The Bilateria or bilaterians are animals with bilateral symmetry as an embryo, i.e. having a left and a right side that are mirror images of each other. This also means they have a head and a tail (anterior-posterior axis) as well as a belly and a back (ventral-dorsal axis). Nearly all are bilaterally symmetrical as adults as well; the most notable exception is the echinoderms, which achieve secondary pentaradial symmetry as adults, but are bilaterally symmetrical during embryonic development. Most animals are bilaterians, excluding sponges, ctenophores, placozoans and cnidarians. For the most part, bilateral embryos are triploblastic, having three germ layers: endoderm, mesoderm, and ectoderm. Except for a few phyla (i.e. flatworms and gnathostomulids), bilaterians have complete digestive tracts with a separate mouth and anus. Some bilaterians lack body cavities (Acoelomata, acoelomates, i.e. Flatworm, Platyhelminthes, Gastrotricha and Gnathostomulida), while others display pri ...
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Ikaria Wariootia
''Ikaria wariootia'' is an early example of a wormlike, bilaterian organism. Its fossils are found in rocks of the Ediacara Member of South Australia that are estimated to be between 560 and 555 million years old. A representative of the Ediacaran biota, ''Ikaria'' lived during the Ediacaran period, roughly 15 million years before the Cambrian, when the Cambrian explosion occurred and where widespread fossil evidence of modern bilaterian taxa appear in the fossil record. Discovery Scott D. Evans, Ian V. Hughes, James G. Gehling, and Mary L. Droser published a paper in the ''Proceedings of the National Academy of Sciences of the United States of America'' on 23 March 2020, describing the finding and identification of ''I. wariootia''. Age The age of Ediacara Member strata are not well-defined through radiometric dating, and are primarily estimated comparatively with other Ediacaran Biota assemblages, likely ranging between approximately 562 Ma and 542 Ma. Brazilian trace f ...
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Xenacoelomorpha
Xenacoelomorpha is a small phylum of bilaterian invertebrate animals, consisting of two sister groups: xenoturbellids and acoelomorphs. This new phylum was named in February 2011 and suggested based on morphological synapomorphies (physical appearances shared by the animals in the clade), which was then confirmed by phylogenomic analyses of molecular data (similarities in the DNA of the animals within the clade). Phylogenetics The clade (groupings of organisms based on their most recent shared/common ancestors) Xenacoelomorpha groups the Acoelomorpha and the genus ''Xenoturbella'', due to molecular studies. Initially this phylum was considered to be a member of the deuterostomes, (meaning during development, as an embryo, the anus develops first and then the mouth), but because of recent transcriptome analysis, it was concluded that phylum Xenacoelomorpha is the sister group (two closest relatives in a phylogenetic tree) to the Nephrozoa, which includes both the protostomes (w ...
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Chaetognatha
The Chaetognatha or chaetognaths (meaning ''bristle-jaws'') are a phylum of predatory marine worms that are a major component of plankton worldwide. Commonly known as arrow worms, about 20% of the known Chaetognatha species are benthic, and can attach to algae and rocks. They are found in all marine waters, from surface tropical waters and shallow tide pools to the deep sea and polar regions. Most chaetognaths are transparent and are torpedo shaped, but some deep-sea species are orange. They range in size from . There are more than 120 modern species assigned to over 20 genera. Despite the limited diversity of species, the number of individuals is large. Arrow worms are usually considered a type of protostome that do not belong to either Ecdysozoa or Lophotrochozoa. Anatomy Chaetognaths are transparent or translucent dart-shaped animals covered by a cuticle. The body is divided into a distinct head, trunk, and tail. There are between four and fourteen hooked, grasping spines ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Ecdysozoa
Ecdysozoa () is a group of protostome animals, including Arthropoda (insects, chelicerata, crustaceans, and myriapods), Nematoda, and several smaller phyla. They were first defined by Aguinaldo ''et al.'' in 1997, based mainly on phylogenetic trees constructed using 18S ribosomal RNA genes. A large study in 2008 by Dunn ''et al.'' strongly supported the Ecdysozoa as a clade, that is, a group consisting of a common ancestor and all its descendants. The group is also supported by morphological characters, and includes all animals that grow by ecdysis, moulting their cuticle. The group was initially contested by a significant minority of biologists. Some argued for groupings based on more traditional taxonomic techniques, while others contested the interpretation of the molecular data. Etymology The name ''Ecdysozoa'' stems etymologically . Characteristics The most notable characteristic shared by ecdysozoans is a three-layered cuticle (four in Tardigrada) composed of orga ...
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Protostomia
Protostomia () is the clade of animals once thought to be characterized by the formation of the organism's mouth before its anus during embryonic development. This nature has since been discovered to be extremely variable among Protostomia's members, although the reverse is typically true of its sister clade, Deuterostomia. Well known examples of protostomes are arthropods, molluscs, annelids, flatworms and nematodes. They are also called schizocoelomates since schizocoely typically occurs in them. Together with the Deuterostomia and Xenacoelomorpha, these form the clade Bilateria, animals with bilateral symmetry, anteroposterior axis and three germ layers. Protostomy In animals at least as complex as earthworms, the first phase in gut development involves the embryo forming a dent on one side (the blastopore) which deepens to become its digestive tube (the archenteron). In the sister-clade, the deuterostomes (), the original dent becomes the anus while the gut event ...
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Spiralia
The Spiralia are a morphologically diverse clade of protostome animals, including within their number the molluscs, annelids, platyhelminths and other taxa. The term ''Spiralia'' is applied to those phyla that exhibit canonical spiral cleavage, a pattern of early development found in most (but not all) members of the Lophotrochozoa. Distribution of spiralian development across phylogeny Members of the molluscs, annelids, platyhelminths and nemerteans have all been shown to exhibit spiral cleavage in its classical form. Other spiralian phyla (rotifers, brachiopods, phoronids, gastrotrichs, and bryozoans) are also said to display a derived form of spiral cleavage in at least a portion of their constituent species, although evidence for this is sparse. Lophotrochozoa within Spiralia Previously, spiral cleavage was thought to be unique to the Spiralia in the strictest sense—animals such as molluscs and annelids which exhibit classical spiral cleavage. The presence of spiral cleav ...
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Ediacaran
The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and the beginning of the Phanerozoic Eon. It is named after the Ediacara Hills of South Australia. The Ediacaran Period's status as an official geological period was ratified in 2004 by the International Union of Geological Sciences (IUGS), making it the first new geological period declared in 120 years. Although the period takes its name from the Ediacara Hills where geologist Reg Sprigg first discovered fossils of the eponymous Ediacaran biota in 1946, the type section is located in the bed of the Enorama Creek within Brachina Gorge in the Flinders Ranges of South Australia, at . The Ediacaran marks the first appearance of widespread multicellular fauna following the end of Snowball Earth glaciation events, the so-called Ediacaran biota, ...
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Deuterostomia
Deuterostomia (; in Greek) are animals typically characterized by their anus forming before their mouth during embryonic development. The group's sister clade is Protostomia, animals whose digestive tract development is more varied. Some examples of deuterostomes include vertebrates (and thus humans), sea stars, and crinoids. In deuterostomy, the developing embryo's first opening (the blastopore) becomes the anus, while the mouth is formed at a different site later on. This was initially the group's distinguishing characteristic, but deuterostomy has since been discovered among protostomes as well. This group is also known as enterocoelomates, because their coelom develops through enterocoely. The three major clades of deuterostomes are Chordata (e.g. vertebrates), Echinodermata (e.g. starfish), and Hemichordata (e.g. acorn worms). Together with Protostomia and their out-group Xenacoelomorpha, these compose the Bilateria, animals with bilateral symmetry and three germ ...
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Nephrozoa
Nephrozoa is a major clade of bilaterians, divided into the protostomes and the deuterostomes, containing almost all animal phyla and over a million extant species. Its sister clade is the Xenacoelomorpha. The Ambulacraria (conventionally deuterostomes) was formerly thought to be sister to the Xenacoelomorpha, forming the Xenambulacraria as basal Deuterostomes, or basal Bilateria invalidating Nephrozoa and Deuterostomes in earlier studies. The coelom, the digestive tract and excretory organs (nephridia), and nerve cords developed in the Nephrozoa. It has been argued that, because protonephridia are only found in protostomes, they cannot be considered a synapomorphy of this group. This would make Nephrozoa an improper name, leaving Eubilateria as this clade's name. Chordates (which include all the vertebrates) are deuterostomes. It seems very likely that the ''Kimberella ''Kimberella'' is an extinct genus of bilaterian known only from rocks of the Ediacaran period. The s ...
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Proarticulata
Proarticulata is a proposed phylum of extinct, bilaterally symmetrical animals known from fossils found in the Ediacaran (Vendian) marine deposits, and dates to approximately . The name comes from the Greek () = "before" and Articulata, i.e. prior to animals with true segmentation such as annelids and arthropods. This phylum was established by Mikhail A. Fedonkin in 1985 for such animals as '' Dickinsonia'', ''Vendia'', ''Cephalonega'', ''Praecambridium'' and currently many other Proarticulata are described (see list). Due to their simplistic morphology, their affinities and mode of life are subject to debate. They are almost universally considered to be metazoans, and due to possessing a clear central axis have been suggested to be stem-bilaterians. In the traditional interpretation, the Proarticulatan body is divided into transverse articulation (division) into isomers as distinct from the transverse articulation segments in annelids and arthropods, as their individual isome ...
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Priapulida
Priapulida (priapulid worms, from Gr. πριάπος, ''priāpos'' 'Priapus' + Lat. ''-ul-'', diminutive), sometimes referred to as penis worms, is a phylum of unsegmented marine worms. The name of the phylum relates to the Greek god of fertility, because their general shape and their extensible spiny introvert (eversible) proboscis may resemble the shape of a human penis. They live in the mud and in comparatively shallow waters up to deep. Some species show a remarkable tolerance for hydrogen sulfide and anoxia. They can be quite abundant in some areas. In an Alaskan bay as many as 85 adult individuals of ''Priapulus caudatus'' per square meter has been recorded, while the density of its larvae can be as high as 58,000 per square meter. Together with Echiura and Sipuncula, they were once placed in the taxon Gephyrea, but consistent morphological and molecular evidence supports their belonging to Ecdysozoa, which also includes arthropods and nematodes. Fossil findings show that ...
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