Allocortex
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Allocortex
The allocortex or heterogenetic cortex, and neocortex are the two types of cerebral cortex in the brain. The allocortex is the much smaller area of cortex taking up just 10 %, the neocortex takes up the remaining 90 %. It is characterized by having just three or four cortical layers, in contrast with the six layers of the neocortex. There are three subtypes of allocortex: the paleocortex, the archicortex, and the periallocortex—a transitional zone between the neocortex and the allocortex. The specific regions of the brain usually described as belonging to the allocortex are the olfactory system, and the hippocampus. Allocortex is termed heterogenetic cortex, because during development it never has the six-layered architecture of homogenetic neocortex. It differs from hetero''typic'' cortex, a type of cerebral cortex, which during prenatal development, passes through a six-layered stage to have fewer layers, such as in Brodmann area 4 that lacks granule cells. Structure The all ...
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Periallocortex
Periallocortex is one of three subtypes of allocortex, the other two subtypes being paleocortex and archicortex. The periallocortex is formed at transition areas where any of the other two subtypes of allocortex borders with the neocortex (which is also called isocortex). Thus, the periallocortex is also subdivided to two subtypes. One subtype is called peripaleocortex, which is formed at borders between paleocortex and neocortex. Areas considered to belong to peripaleocortex are for example anterior insular cortex. Another subtype of periallocortex is called periarchicortex. It is formed at borders between archicortex and neocortex. Areas considered to belong to periarchicortex include entorhinal cortex, perirhinal cortex, presubiculum, parasubiculum, retrosplenial cortex, subcallosal area and subgenual area. No one allocortex or even periallocortex area borders, contacts or transitions immediately to the so-called true isocortex. Instead, they border and slowly transition to an ...
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Periallocortex
Periallocortex is one of three subtypes of allocortex, the other two subtypes being paleocortex and archicortex. The periallocortex is formed at transition areas where any of the other two subtypes of allocortex borders with the neocortex (which is also called isocortex). Thus, the periallocortex is also subdivided to two subtypes. One subtype is called peripaleocortex, which is formed at borders between paleocortex and neocortex. Areas considered to belong to peripaleocortex are for example anterior insular cortex. Another subtype of periallocortex is called periarchicortex. It is formed at borders between archicortex and neocortex. Areas considered to belong to periarchicortex include entorhinal cortex, perirhinal cortex, presubiculum, parasubiculum, retrosplenial cortex, subcallosal area and subgenual area. No one allocortex or even periallocortex area borders, contacts or transitions immediately to the so-called true isocortex. Instead, they border and slowly transition to an ...
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Neocortex
The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, spatial reasoning and language. The neocortex is further subdivided into the true isocortex and the proisocortex. In the human brain, the neocortex is the largest part of the cerebral cortex (the outer layer of the cerebrum). The neocortex makes up the largest part of the cerebral cortex, with the allocortex making up the rest. The neocortex is made up of six layers, labelled from the outermost inwards, I to VI. Etymology The term is from ''cortex'', Latin, " bark" or "rind", combined with ''neo-'', Greek, "new". ''Neopallium'' is a similar hybrid, from Latin ''pallium'', "cloak". ''Isocortex'' and ''allocortex'' are hybrids with Greek ''isos'', "same", and ''allos'', "other". Anatomy The neocortex is the most developed in its organisat ...
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Piriform Cortex
The piriform cortex, or pyriform cortex, is a region in the brain, part of the rhinencephalon situated in the cerebrum. The function of the piriform cortex relates to the sense of smell. Structure The piriform cortex is part of the rhinencephalon situated in the cerebrum. In human anatomy, the piriform cortex has been described as consisting of the cortical amygdala, uncus, and anterior parahippocampal gyrus. More specifically, the human piriform cortex is located between the insula and the temporal lobe, anteriorly and laterally of the amygdala.Howard, J. D., Plailly, J., Grueschow, M., Haynes, J. D., & Gottfried, J. A. (2009). Odor quality coding and categorization in human posterior piriform cortex. Nature neuroscience, 12(7), 932-938. Supplementary material, p.4 Function The function of the piriform cortex relates to olfaction, which is the perception of smell. This has been particularly shown in humans for the posterior piriform cortex. The piriform cortex in rodents and ...
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Phylogenetics
In biology, phylogenetics (; from Greek language, Greek wikt:φυλή, φυλή/wikt:φῦλον, φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, Protein, protein Amino acid, amino acid sequences, or Morphology (biology), morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An un ...
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Brodmann Area 25
Brodmann area 25 (BA25) is the subgenual area, area subgenualis or subgenual cingulatea area in the cerebral cortex of the brain and delineated based on its cytoarchitectonic characteristics. It is the 25th "Brodmann area" defined by Korbinian Brodmann (thus its name). BA25 is located in the cingulate region as a narrow band in the caudal portion of the subcallosal area adjacent to the paraterminal gyrus. The posterior parolfactory sulcus separates the paraterminal gyrus from BA25. Rostrally it is bound by the prefrontal area 11 of Brodmann. History Brodmann described this area as it is labeled now in 1909. Originally in 1905, Brodmann labeled the area as part of area 24. In 1909, he divided the area into area 24 and 25. Function This region is extremely rich in serotonin transporters and is considered as a governor for a vast network involving areas like hypothalamus and brain stem, which influences changes in appetite and sleep; the amygdala and insula, which affect the ...
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Indusium Griseum
The indusium griseum, (supracallosal gyrus, gyrus epicallosus) consists of a thin membranous layer of grey matter in contact with the upper surface of the corpus callosum and continuous laterally with the grey matter of the cingulate cortex and inferiorly with the hippocampus. It is vestigial in humans and is a remnant of the former position of the hippocampus in lower animals. On either side of the midline of the indusium griseum are two ridges formed by bands of longitudinally directed fibers known as the medial and lateral longitudinal striae. The indusium griseum is prolonged around the splenium of the corpus callosum as a delicate layer, the fasciolar gyrus, which is continuous below with the surface of the dentate gyrus. Toward the genu of the corpus callosum it curves down along the rostrum to form the subcallosal gyrus The subcallosal gyrus (paraterminal gyrus, peduncle of the corpus callosum) is a narrow lamina on the medial surface of the hemisphere in front of the lam ...
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Retrosplenial Cortex
The retrosplenial cortex (RSC) is a cortical area in the brain comprising Brodmann areas 29 and 30. It is secondary association cortex, making connections with numerous other brain regions. The region's name refers to its anatomical location immediately behind the splenium of the corpus callosum in primates, although in rodents it is located more towards the brain surface and is relatively larger. Its function is currently not well understood, but its location close to visual areas and also to the hippocampal spatial/memory system suggest it may have a role in mediating between perceptual and memory functions, particularly in the spatial domain. However, its exact contribution to either space or memory processing has been hard to pin down. Anatomy There is large variation in the region's size across different species. In humans it comprises roughly 0.3% of the entire cortical surface whereas in rabbits it is at least 10% and in rats it extends for more than half the cerebrum do ...
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Entorhinal Cortex
The entorhinal cortex (EC) is an area of the brain's allocortex, located in the medial temporal lobe, whose functions include being a widespread network hub for memory, navigation, and the perception of time.Integrating time from experience in the lateral entorhinal cortex Albert Tsao, Jørgen Sugar, Li Lu, Cheng Wang, James J. Knierim, May-Britt Moser & Edvard I. Moser Naturevolume 561, pages57–62 (2018) The EC is the main interface between the hippocampus and neocortex. The EC-hippocampus system plays an important role in declarative (autobiographical/episodic/semantic) memories and in particular spatial memories including memory formation, memory consolidation, and memory optimization in sleep. The EC is also responsible for the pre-processing (familiarity) of the input signals in the reflex nictitating membrane response of classical trace conditioning; the association of impulses from the eye and the ear occurs in the entorhinal cortex. Structure In rodents, the EC ...
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Periarchicortex
Periarchicortex is one of two subtypes of periallocortex, the other being peripaleocortex. It is formed at borders between archicortex (a subtype of allocortex) and isocortex and shows slow histological transition from the four-layered structure typical for archicortex to the six-layered structure typical for isocortex. Cortical areas that are generally considered to belong to periarchicortex, include presubiculum, parasubiculum, entorhinal cortex, perirhinal cortex, retrosplenial cortex, periarchicortical part of cingulate cortex, posterior part of subcallosal area, and subgenual area. Periarchicortex does not contact immediately at borders with the true isocortex and does not transit directly into it. Instead, another transitional area from the isocortex side, called proisocortex Proisocortex or pro-isocortex is one of two subtypes of cortical areas in the areas belonging to the neocortex. The other subtype is termed the true isocortex. Proisocortical areas are transitional area ...
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Dentate Gyrus
The dentate gyrus (DG) is part of the hippocampal formation in the temporal lobe of the brain, which also includes the hippocampus and the subiculum. The dentate gyrus is part of the hippocampal trisynaptic circuit and is thought to contribute to the formation of new episodic memories, the spontaneous exploration of novel environments and other functions. It is notable as being one of a select few brain structures known to have significant rates of adult neurogenesis in many species of mammals, from rodents to primates. Other sites of adult neurogenesis include the subventricular zone, the striatum and the cerebellum. However, whether significant neurogenesis exists in the adult human dentate gyrus has been a matter of debate. 2019 evidence has shown that adult neurogenesis does take place in the subventricular zone and in the subgranular zone of the dentate gyrus. Structure The dentate gyrus, like the hippocampus, consists of three distinct layers: an outer molecular layer ...
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Cerebral Cortex
The cerebral cortex, also known as the cerebral mantle, is the outer layer of neural tissue of the cerebrum of the brain in humans and other mammals. The cerebral cortex mostly consists of the six-layered neocortex, with just 10% consisting of allocortex. It is separated into two cortices, by the longitudinal fissure that divides the cerebrum into the left and right cerebral hemispheres. The two hemispheres are joined beneath the cortex by the corpus callosum. The cerebral cortex is the largest site of neural integration in the central nervous system. It plays a key role in attention, perception, awareness, thought, memory, language, and consciousness. The cerebral cortex is part of the brain responsible for cognition. In most mammals, apart from small mammals that have small brains, the cerebral cortex is folded, providing a greater surface area in the confined volume of the cranium. Apart from minimising brain and cranial volume, cortical folding is crucial for the brain ...
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