Amphioxus
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Amphioxus
The lancelets ( or ), also known as amphioxi (singular: amphioxus ), consist of some 30 to 35 species of "fish-like" benthic filter feeding chordates in the order Amphioxiformes. They are the modern representatives of the subphylum Cephalochordata. Lancelets closely resemble 530-million-year-old ''Pikaia'', fossils of which are known from the Burgess Shale. Zoologists are interested in them because they provide evolutionary insight into the origins of vertebrates. Lancelets contain many organs and organ systems that are closely related to those of modern fish, but in more primitive form. Therefore, they provide a number of examples of possible evolutionary exaptation. For example, the gill-slits of lancelets are used for feeding only, and not for respiration. The circulatory system carries food throughout their body, but does not have red blood cells or hemoglobin for transporting oxygen. Lancelet genomes hold clues about the early evolution of vertebrates: by comparing genes from ...
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Linda Holland
Linda Zimmerman Holland is a research biologist at Scripps Institution of Oceanography known for her work examining the evolution of vertebrates. Education and career Holland has a B.A. (1962) and an M.A. (1964) from Stanford University. She worked as a research associate at the University of California, San Diego, Scripps Clinic, and Scripps Institution of Oceanography in the period from 1970 until 1998. She earned her Ph.D. from the University of California, San Diego in 2001. She started as a research biologist at Scripps in 1998, and is emeritus as of 2022. Linda Holland has described in detail some of the early obstacles she faced as a woman scientist starting out in academe in the 1960s and 1970s, Research Holland's early research examined anatomical structures in purple sea urchins, and a protein involved in clotting, Von Willebrand factor. She went on to examine reproduction in sea urchins, salps, and amphioxus, known as lancelet. Holland began collecting amphioxus in ...
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Branchiostoma Lanceolatum
''Branchiostoma lanceolatum'', the European lancelet or Mediterranean amphioxus, is a lancelet in the subphylum Cephalochordata. It is a marine invertebrate with a notochord but no backbone and is used as a model organism to study the evolutionary development of vertebrates. Anatomy ''Branchiostoma lanceolatum'' has an elongated body, flattened laterally and pointed at both ends. A stiffening rod of tightly packed cells, the notochord, extends the whole length of the body. Unlike vertebrates, the notochord persists in the adult, in form of simple dorsal neural tube slightly thickened in the anterior part (the cerebral vesicle). Above it is a nerve cord with a single frontal eye. The mouth is on the underside of the body and is surrounded by a tuft of 20 or 30 cirri or slender sensory appendages. The gut runs just below the notochord from the mouth to the anus, in front of the tail. There is a flap-like, vertical fin surrounding the pointed tail. Gas exchange takes place as w ...
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Chordates
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cephalochordates. These CSIs prov ...
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Pikaia
''Pikaia gracilens'' is an extinct, primitive chordate animal known from the Middle Cambrian Burgess Shale of British Columbia. Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became the "most famous early chordate fossil," or "famously known as the earliest" chordates. It is estimated to have lived during the latter period of the Cambrian explosion. Since it initial discovery, more than a hundred specimens have been recovered. The body structure resembles that of the lancelet and perhaps it swam much like an eel. A notochord and myomeres (segmented blocks of skeletal muscles) span the entire length of the body, and are considered as the defining signatures of chordate characters. Its primitive feature is indicated by the body covering, a cuticle, which is characteristic of invertebrates and some protochordates. The exact phylogenetic position is unclear. Proposed affinities include ...
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Lampreys
Lampreys (sometimes inaccurately called lamprey eels) are an ancient extant lineage of jawless fish of the order Petromyzontiformes , placed in the superclass Cyclostomata. The adult lamprey may be characterized by a toothed, funnel-like sucking mouth. The common name "lamprey" is probably derived from Latin , which may mean "stone licker" ( "to lick" + "stone"), though the etymology is uncertain. ''Lamprey'' is sometimes seen for the plural form. There are about 38 known extant species of lampreys and five known extinct species. Parasitic carnivorous species are the most well-known, and feed by boring into the flesh of other fish to suck their blood; but only 18 species of lampreys engage in this micropredatory lifestyle. Of the 18 carnivorous species, nine migrate from saltwater to freshwater to breed (some of them also have freshwater populations), and nine live exclusively in freshwater. All non-carnivorous forms are freshwater species. Adults of the non-carnivorou ...
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Cephalochordata
A cephalochordate (from Greek language, Greek: κεφαλή ''kephalé'', "head" and χορδή ''khordé'', "chord") is an animal in the chordate subphylum, Cephalochordata. They are commonly called Lancelet, lancelets. Cephalochordates possess 5 synapomorphies, or primary characteristics, that all chordates have at some point during their larval or adulthood stages. These 5 synapomorphies include a notochord, dorsal hollow nerve cord, endostyle, Pharyngeal slit, pharyngeal slits, and a post-anal tail (see chordate for descriptions of each). The fine structure of the cephalochordate notochord is best known for the Bahamas lancelet, ''Asymmetron lucayanum''. Cephalochordates are represented in modern oceans by the Amphioxiformes and are commonly found in warm temperate and tropical seas worldwide. With the presence of a notochord, adult amphioxus are able to swim and tolerate the tides of coastal environments, but they are most likely to be found within the sediment of these commun ...
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Benthos
Benthos (), also known as benthon, is the community of organisms that live on, in, or near the bottom of a sea, river, lake, or stream, also known as the benthic zone.Benthos
from the Census of Antarctic Marine Life website
This community lives in or near marine or freshwater sedimentary environments, from s along the , out to the , and t ...
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Gonochorism
In biology, gonochorism is a sexual system where there are only two Sex, sexes and each individual organism is either male or female. The term gonochorism is usually applied in animal species, the vast majority of which are gonochoric. Gonochorism contrasts with Hermaphrodite, simultaneous hermaphroditism but it may be hard to tell if a species is gonochoric or sequentially hermaphroditic. (e.g. Parrotfish, ''Patella ferruginea''). However, in gonochoric species individuals remain either male or female throughout their lives. Species that reproduce by Thelytokous, thelytokous parthenogenesis and do not have males can still be classified as gonochoric. Terminology The term is derived from Greek language, Greek (''gone'', generation) + (''chorizein,'' to separate). The term gonochorism originally came from German gonochorismus. Gonochorism is also referred to as unisexualism or gonochory. Evolution Gonochorism has Convergent evolution, evolved independently multiple time ...
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Fertilisation
Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Processes such as insemination or pollination which happen before the fusion of gametes are also sometimes informally called fertilisation. The cycle of fertilisation and development of new individuals is called sexual reproduction. During double fertilisation in angiosperms the haploid male gamete combines with two haploid polar nuclei to form a triploid primary endosperm nucleus by the process of vegetative fertilisation. History In Antiquity, Aristotle conceived the formation of new individuals through fusion of male and female fluids, with form and function emerging gradually, in a mode called by him as epigenetic. In 1784, Spallanzani established the need of interaction between the female's ovum and male's sperm to form a zygote in frog ...
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Spawn (biology)
Spawn is the Egg cell, eggs and Spermatozoa, sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of releasing the eggs and sperm, and the act of both sexes is called spawning. Most aquatic animals, except for aquatic mammals and marine reptile, reptiles, reproduce through the process of spawning. Spawn consists of the reproductive cells (gametes) of many aquatic animals, some of which will become fertilized and produce offspring. The process of spawning typically involves females releasing Ovum, ova (unfertilized eggs) into the water, often in large quantities, while males simultaneously or sequentially release spermatozoa (milt) to fertilize the eggs. Most fish reproduce by spawning, as do most other aquatic animals, including crustaceans such as crabs and shrimps, molluscs such as oysters and squid, echinoderms such as sea urchins and sea cucumbers, amphibians such as frogs and newts, aquatic insects such as mayflies and mos ...
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Limax
''Limax'' is a genus of air-breathing land slugs in the terrestrial pulmonate gastropod mollusk family Limacidae. The generic name ''Limax'' literally means "slug". Some species, such as the leopard slug (''L. maximus'') and the tawny garden slug (''Limacus flavus''), are considered garden pests.Ebeling. WChapter 12: Miscellaneous Pests. ''Urban Entomology''. Department of Entomology. UC Riverside. 2002. The genus ''Limax'' ''sensu stricto'' is probably monophyletic.Klee, B. 200Towards a phylogeny of ''Limax'' (Gastropoda: Stylommatophora)''. ''The Malacologist''. Molluscan Forum 2006. Distribution This genus is native to Europe, but at least one species (''L. maximus'') has been introduced into North America. Species The genus ''Limax'' includes at least 33 species:Species in genus ''Lima ...
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Peter Simon Pallas
Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussian zoologist and botanist who worked in Russia between 1767 and 1810. Life and work Peter Simon Pallas was born in Berlin, the son of Professor of Surgery Simon Pallas. He studied with private tutors and took an interest in natural history, later attending the University of Halle and the University of Göttingen. In 1760, he moved to the University of Leiden and passed his doctor's degree at the age of 19. Pallas travelled throughout the Netherlands and to London, improving his medical and surgical knowledge. He then settled at The Hague, and his new system of animal classification was praised by Georges Cuvier. Pallas wrote ''Miscellanea Zoologica'' (1766), which included descriptions of several vertebrates new to science which he had discovered in the Dutch museum collections. A planned voyage to southern Africa and the East Indies fell through when his father reca ...
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