Wulagasaurus
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Wulagasaurus
''Wulagasaurus'' (meaning "Wulaga lizard", in reference to the discovery locality) is a genus of saurolophine hadrosaurid dinosaur from the Late Cretaceous of Heilongjiang, China. Its remains were found in a bonebed in the middle Maastrichtian-age Yuliangze Formation, dated to 69 million years ago.Godefroit, P., Lauters, P., Van Itterbeeck, J., Bolotsky, Y. and Bolotsky, I.Y. (2011). "Recent advances on study of hadrosaurid dinosaurs in Heilongjiang (Amur) River area between China and Russia." ''Global Geology'', 2011(3). This bonebed is otherwise dominated by fossils of the lambeosaurine hadrosaurid (hollow-crested duckbill) '' Sahaliyania''. ''Wulagasaurus'' was named by Pascal Godefroit and colleagues in 2008. Only partial remains are known at this time. It is one of several hadrosaurids from the Amur River region named since 2000. The type and only species to date is ''W. dongi'', named in honor of Chinese paleontologist Dong Zhiming. In 2010 Gregory S. Paul estimated its s ...
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Wulagasaurus Revision
''Wulagasaurus'' (meaning "Wulaga lizard", in reference to the discovery locality) is a genus of saurolophine hadrosaurid dinosaur from the Late Cretaceous of Heilongjiang, China. Its remains were found in a bonebed in the middle Maastrichtian-age Yuliangze Formation, dated to 69 million years ago.Godefroit, P., Lauters, P., Van Itterbeeck, J., Bolotsky, Y. and Bolotsky, I.Y. (2011). "Recent advances on study of hadrosaurid dinosaurs in Heilongjiang (Amur) River area between China and Russia." ''Global Geology'', 2011(3). This bonebed is otherwise dominated by fossils of the lambeosaurine hadrosaurid (hollow-crested duckbill) ''Sahaliyania''. ''Wulagasaurus'' was named by Pascal Godefroit and colleagues in 2008. Only partial remains are known at this time. It is one of several hadrosaurids from the Amur River region named since 2000. The type and only species to date is ''W. dongi'', named in honor of Chinese paleontologist Dong Zhiming. In 2010 Gregory S. Paul estimated its si ...
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Yuliangze Formation
The Yuliangze Formation, or Yuliangzi Formation, is a geological formation in Heilongjiang, China whose strata date back to the early-middle Maastrichtian. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Cretaceous, Asia)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 593-600. . Vertebrate paleofauna * '' Charonosaurus jiayinensis'' - "Partial skull and partial, fragmentary postcranial elements.""Table 20.1," in Weishampel, et al. (2004). Page 441. * ''Mandschurosaurus amurensis'' (Hadrosaurinae indet.) * '' Sahaliyania elunchunorum'' ( Hadrosauridae indet.) * ''Tarbosaurus bataar'' *'' Saurolophus krystofovici'' * '' Wulagasaurus dongi'' See also * List of dinosaur-bearing rock formations This list of dinosaur-bearing rock formations is a list of geologic formations in which dinosaur fos ...
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Saurolophinae
Saurolophinae is a subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. However, the name Hadrosaurinae is based on the genus ''Hadrosaurus'' which was found in more recent studies to be more primitive than either lambeosaurines or other traditional "hadrosaurines", like ''Edmontosaurus'' and '' Saurolophus''. As a result of this, the name Hadrosaurinae was dropped or restricted to ''Hadrosaurus'' alone, and the subfamily comprising the traditional "hadrosaurines" was renamed the Saurolophinae. Recent phylogenetic work by Hai Xing indicates that ''Hadrosaurus'' is placed within the monophyletic group containing all non-lambeosaurine hadrosaurids. Under this view, the traditional Hadrosaurinae is resurrected, with the Hadrosauridae being divided into two clades: Hadrosaurinae and Lambeosaurinae. Saurolophinae was first defined as ...
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Hadrosaurid
Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includes genera such as ''Edmontosaurus'' and '' Parasaurolophus'', was a common group of herbivores during the Late Cretaceous Period. Hadrosaurids are descendants of the Upper Jurassic/Lower Cretaceous iguanodontian dinosaurs and had a similar body layout. Hadrosaurs were among the most dominant herbivores during the Late Cretaceous in Asia and North America, and during the close of the Cretaceous several lineages dispersed into Europe, Africa, South America and Antarctica. Like other ornithischians, hadrosaurids had a predentary bone and a pubic bone which was positioned backwards in the pelvis. Unlike more primitive iguanodonts, the teeth of hadrosaurids are stacked into complex structures known as dental batteries, which acted as effectiv ...
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Sahaliyania
''Sahaliyania'' (from "black" in Manchu, a reference to the Amur/ Heilongjiang River) is a genus of lambeosaurine hadrosaurid dinosaur (crested duckbilled dinosaur) from the Late Cretaceous of Heilongjiang, China. Discovery Its remains were found in a bonebed in the Maastrichtian-age Yuliangze Formation, alongside rarer remains of the hadrosaurine hadrosaurid (flat-headed duckbill) ''Wulagasaurus''. ''Sahaliyania'' was named by Pascal Godefroit and colleagues in 2008. It is one of several hadrosaurids from the Amur River region named since 2000. The type and only species to date is ''S. elunchunorum'', named in honor of the Elunchun people. ''Sahaliyania'' is based on GMH W453, a partial skull. Godefroit and colleagues assigned numerous other fossils from the bonebed to their new genus, representing much of the skull, pectoral girdle, upper arm, and pelvis. It can be distinguished from other hadrosaurids by a variety of anatomical details. Godefroit and colleagues perf ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Cladistics
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Autapomorphy
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to the focal taxon (which may be a species, family or in general any clade). It can therefore be considered an apomorphy in relation to a single taxon. The word ''autapomorphy'', first introduced in 1950 by German entomologist Willi Hennig, is derived from the Greek words αὐτός, ''autos'' "self"; ἀπό, ''apo'' "away from"; and μορφή, ''morphḗ'' = "shape". Discussion Because autapomorphies are only present in a single taxon, they do not convey information about relationship. Therefore, autapomorphies are not useful to infer phylogenetic relationships. However, autapomorphy, like synapomorphy and plesiomorphy is a relative concept depending on the taxon in question. An autapomorphy at a given level may well be a synapomorphy at ...
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Ischium
The ischium () forms the lower and back region of the (''os coxae''). Situated below the ilium and behind the pubis, it is one of three regions whose fusion creates the . The superior portion of this region forms approximately one-third of the acetabulum.


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Humerus
The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a rounded head, a narrow neck, and two short processes (tubercles, sometimes called tuberosities). The body is cylindrical in its upper portion, and more prismatic below. The lower extremity consists of 2 epicondyles, 2 processes (trochlea & capitulum), and 3 fossae (radial fossa, coronoid fossa, and olecranon fossa). As well as its true anatomical neck, the constriction below the greater and lesser tubercles of the humerus is referred to as its surgical neck due to its tendency to fracture, thus often becoming the focus of surgeons. Etymology The word "humerus" is derived from la, humerus, umerus meaning upper arm, shoulder, and is linguistically related to Gothic ''ams'' shoulder and Greek ''ōmos''. Structure Upper extremity The upper or pr ...
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Geological Museum Of Heilongjiang
Geology () is a branch of natural science concerned with Earth and other astronomical objects, the features or rocks of which it is composed, and the processes by which they change over time. Modern geology significantly overlaps all other Earth sciences, including hydrology, and so is treated as one major aspect of integrated Earth system science and planetary science. Geology describes the structure of the Earth on and beneath its surface, and the processes that have shaped that structure. It also provides tools to determine the relative and absolute ages of rocks found in a given location, and also to describe the histories of those rocks. By combining these tools, geologists are able to chronicle the geological history of the Earth as a whole, and also to demonstrate the age of the Earth. Geology provides the primary evidence for plate tectonics, the evolutionary history of life, and the Earth's past climates. Geologists broadly study the properties and processes of Earth ...
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Sternum
The sternum or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major blood vessels from injury. Shaped roughly like a necktie, it is one of the largest and longest flat bones of the body. Its three regions are the manubrium, the body, and the xiphoid process. The word "sternum" originates from the Ancient Greek στέρνον (stérnon), meaning "chest". Structure The sternum is a narrow, flat bone, forming the middle portion of the front of the chest. The top of the sternum supports the clavicles (collarbones) and its edges join with the costal cartilages of the first two pairs of ribs. The inner surface of the sternum is also the attachment of the sternopericardial ligaments. Its top is also connected to the sternocleidomastoid muscle. The sternum consists of three main parts, listed from the top: * Manubrium * Body (gladiolus) * ...
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