Wing Coupling (Lepidoptera Anatomy)
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Wing Coupling (Lepidoptera Anatomy)
Some four-winged insect orders, such as the Lepidoptera, have developed a wide variety of morphological wing coupling mechanisms in the imago which render these taxa as "functionally dipterous" (effectively two-winged) for efficient insect flight. All but the most basal forms exhibit this wing coupling. The mechanisms are of three different types - jugal, frenulo-retinacular and amplexiform. Subsec. "Wing coupling". Pp 56-60. Jugal wing coupling The more primitive groups of moth have an enlarged lobe-like area near the basal posterior margin, i.e. at the base of the forewing, called ''jugum'', that folds under the hindwing in flight. Frenulo-retinacular wing coupling Other groups of moth have a frenulum on the hindwing that hooks under a retinaculum on the forewing. The retinaculum is a hook or tuft on the underside of the forewing of some moths. Along with the frenulum, a spine at the base of the forward or costal edge of the hindwing, it forms a coupling mechanism for the f ...
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Lepidoptera Psychidae Oiketicus Freno O Frenulo
Lepidoptera ( ) is an order of insects that includes butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 percent of the total described species of living organisms. It is one of the most widespread and widely recognizable insect orders in the world. The Lepidoptera show many variations of the basic body structure that have evolved to gain advantages in lifestyle and distribution. Recent estimates suggest the order may have more species than earlier thought, and is among the four most speciose orders, along with the Hymenoptera, Diptera, and Coleoptera. Lepidopteran species are characterized by more than three derived features. The most apparent is the presence of scales that cover the bodies, wings, and a proboscis. The scales are modified, flattened "hairs", and give butterflies and moths their wide variety of colors and patterns. Almost all species have some form of memb ...
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Lepidoptera
Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic rank, superfamilies, 10 percent of the total described species of living organisms. It is one of the most widespread and widely recognizable insect orders in the world. The Lepidoptera show many variations of the basic body structure that have evolved to gain advantages in lifestyle and distribution. Recent estimates suggest the order may have more species than earlier thought, and is among the four most wikt:speciose, speciose orders, along with the Hymenoptera, fly, Diptera, and beetle, Coleoptera. Lepidopteran species are characterized by more than three derived features. The most apparent is the presence of scale (anatomy), scales that cover the torso, bodies, wings, and a proboscis. The scales are modified, flattened "hairs", and give ...
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Imago
In biology, the imago (Latin for "image") is the last stage an insect attains during its metamorphosis, its process of growth and development; it is also called the imaginal stage, the stage in which the insect attains maturity. It follows the final ecdysis of the immature instars.Carpenter, Geo. H., The Life-Story of Insects. Cambridge University Press 1913. May be downloaded from: https://www.gutenberg.org/ebooks/16410 or https://archive.org/details/thelifestoryofin16410gut In a member of the Ametabola or Hemimetabola, in which metamorphosis is "incomplete", the final ecdysis follows the last immature or '' nymphal'' stage. In members of the Holometabola, in which there is a pupal stage, the final ecdysis follows emergence from the pupa, after which the metamorphosis is complete, although there is a prolonged period of maturation in some species. The imago is the only stage during which the insect is sexually mature and, if it is a winged species, has functional wings. The i ...
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Insect Flight
Insects are the only group of invertebrates that have evolved wings and flight. Insects first flew in the Carboniferous, some 350 to 400 million years ago, making them the first animals to evolve flight. Wings may have evolved from appendages on the sides of existing limbs, which already had nerves, joints, and muscles used for other purposes. These may initially have been used for sailing on water, or to slow the rate of descent when gliding. Two insect groups, the dragonflies and mayflies, have flight muscles attached directly to the wings. In other winged insects, flight muscles attach to the thorax, which make it oscillate in order to induce the wings to beat. Of these insects, some (flies and some beetles) achieve very high wingbeat frequencies through the evolution of an "asynchronous" nervous system, in which the thorax oscillates faster than the rate of nerve impulses. Not all insects are capable of flight. A number of apterous insects have secondarily lost their w ...
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Moth
Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of which have yet to be described. Most species of moth are nocturnal, but there are also crepuscular and diurnal species. Differences between butterflies and moths While the butterflies form a monophyletic group, the moths, comprising the rest of the Lepidoptera, do not. Many attempts have been made to group the superfamilies of the Lepidoptera into natural groups, most of which fail because one of the two groups is not monophyletic: Microlepidoptera and Macrolepidoptera, Heterocera and Rhopalocera, Jugatae and Frenatae, Monotrysia and Ditrysia.Scoble, MJ 1995. The Lepidoptera: Form, function and diversity. Oxford, UK: Oxford University Press; 404 p. Although the rules for distinguishing moths from butterflies are not well establishe ...
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Butterflies
Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises the large superfamily (zoology), superfamily Papilionoidea, which contains at least one former group, the skippers (formerly the superfamily "Hesperioidea"), and the most recent analyses suggest it also contains the moth-butterflies (formerly the superfamily "Hedyloidea"). Butterfly fossils date to the Paleocene, about 56 million years ago. Butterflies have a four-stage life cycle, as like most insects they undergo Holometabolism, complete metamorphosis. Winged adults lay eggs on the food plant on which their larvae, known as caterpillars, will feed. The caterpillars grow, sometimes very rapidly, and when fully developed, pupate in a chrysalis. When metamorphosis is complete, the pupal skin splits, the adult insect climbs o ...
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Skipper (butterfly)
Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy places the family in the superfamily Papilionoidea, the butterflies. They are named for their quick, darting flight habits. Most have their antenna tips modified into narrow, hook-like projections. Moreover, skippers mostly have an absence of wing-coupling structure available in most moths. More than 3500 species of skippers are recognized, and they occur worldwide, but with the greatest diversity in the Neotropical regions of Central and South America.Ackery et al. (1999) Description and systematics Traditionally, the Hesperiidae were placed in a monotypic superfamily Hesperioidea, because they are morphologically distinct from other Rhopalocera (butterflies), which mostly belong to the typical butterfly superfamily Papilionoidea. The ...
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Bombycoidea
Bombycoidea is a superfamily of moths. It contains the silk moths, emperor moths, sphinx moths, and relatives. The Lasiocampoidea The Lasiocampidae are a family of moths also known as eggars, tent caterpillars, snout moths (although this also refers to the Pyralidae), or lappet moths. Over 2,000 species occur worldwide, and probably not all have been named or studied. It ... are close relatives and were historically sometimes merged in this group. After many years of debate and shifting taxonomies, the most recent classifications treat the superfamily as containing 10 constituent families. Their larvae often exhibit horns.''Firefly Encyclopedia of Insects and Spiders'', edited by Christopher O'Toole, , 2002 References Lepidoptera superfamilies Macroheterocera {{Bombycoidea-stub ...
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Sphingidae
The Sphingidae are a family of moths (Lepidoptera) called sphinx moths, also colloquially known as hawk moths, with many of their caterpillars known as “hornworms”; it includes about 1,450 species. It is best represented in the tropics, but species are found in every region.Scoble, Malcolm J. (1995): ''The Lepidoptera: Form, Function and Diversity'' (2nd edition). Oxford University Press & Natural History Museum London. They are moderate to large in size and are distinguished among moths for their agile and sustained flying ability, similar enough to that of hummingbirds as to be reliably mistaken for them. Their narrow wings and streamlined abdomens are adaptations for rapid flight. The family was named by French zoologist Pierre André Latreille in 1802. Some hawk moths, such as the hummingbird hawk-moth or the white-lined sphinx, hover in midair while they feed on nectar from flowers, so are sometimes mistaken for hummingbirds. This hovering capability is only known to ...
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