Walcottidiscus
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Walcottidiscus
''Walcottidiscus'' is a genus of pentagonally shaped echinoderms known from the Middle Cambrian Burgess Shale. 16 specimens of ''Walcottidiscus'' are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community. The genus is thought to be the earliest known, more or less undisputed . The pentamerous ''
Arkarua ''Arkarua adami'' is a small, Precambrian disk-like fossil with a raised center, a number of radial ridges on the rim, and a five-pointed central depression marked with radial lines of five small dots from the middle of the disk center. Fossils ...
'' is older, dating from the late
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Edrioasteroidea
Edrioasteroidea is an extinct class of echinoderms. The living animal would have resembled a pentamerously symmetrical disc or cushion. They were obligate encrusters and attached themselves to inorganic or biologic hard substrates (frequently hardgrounds or brachiopods). A 507 million years old species, Totiglobus spencensis, is actually the first known echinoderm adapted to live on a hard surface after the soft microbial mats that covered the seafloor were destroyed in the Cambrian substrate revolution. The oldest undisputed fossils of Edrioasteroidea are known from Cambrian ( Stage 3, about 515-520 Ma ago) of Laurentia and are among the oldest known fossils of echinoderms. Some authors propose that an enigmatic Ediacaran (about 600 Ma) organism '' Arkarua'' is also an edrioasteroid, but this interpretation did not gain wide acceptance. Last edrioasteroids are known from Permian (Late Kungurian, about 270-280 Ma). Anatomy The body plan for this class was simple: a main body ( ...
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Edrioasteroid
Edrioasteroidea is an extinct class (biology), class of echinoderms. The living animal would have resembled a pentamerously symmetrical disc or cushion. They were obligate encrusters and attached themselves to inorganic or biologic hard substrates (frequently hardgrounds or brachiopods). A 507 million years old species, Totiglobus spencensis, is actually the first known echinoderm adapted to live on a hard surface after the soft microbial mats that covered the seafloor were destroyed in the Cambrian substrate revolution. The oldest undisputed fossils of Edrioasteroidea are known from Cambrian (Cambrian Stage 3, Stage 3, about 515-520 Megaannum, Ma ago) of Laurentia and are among the oldest known fossils of echinoderms. Some authors propose that an enigmatic Ediacaran (about 600 Ma) organism ''Arkarua'' is also an edrioasteroid, but this interpretation did not gain wide acceptance. Last edrioasteroids are known from Permian (Late Kungurian, about 270-280 Ma). Anatomy The body plan ...
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Pentagon
In geometry, a pentagon (from the Greek πέντε ''pente'' meaning ''five'' and γωνία ''gonia'' meaning ''angle'') is any five-sided polygon or 5-gon. The sum of the internal angles in a simple pentagon is 540°. A pentagon may be simple or self-intersecting. A self-intersecting ''regular pentagon'' (or ''star pentagon'') is called a pentagram. Regular pentagons A '' regular pentagon'' has Schläfli symbol and interior angles of 108°. A '' regular pentagon'' has five lines of reflectional symmetry, and rotational symmetry of order 5 (through 72°, 144°, 216° and 288°). The diagonals of a convex regular pentagon are in the golden ratio to its sides. Given its side length t, its height H (distance from one side to the opposite vertex), width W (distance between two farthest separated points, which equals the diagonal length D) and circumradius R are given by: :\begin H &= \frac~t \approx 1.539~t, \\ W= D &= \frac~t\approx 1.618~t, \\ W &= \sqr ...
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Echinoderm
An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea lilies or "stone lilies". Adult echinoderms are found on the sea bed at every ocean depth, from the intertidal zone to the abyssal zone. The phylum contains about 7,000 living species, making it the second-largest grouping of deuterostomes, after the chordates. Echinoderms are the largest entirely marine phylum. The first definitive echinoderms appeared near the start of the Cambrian. The echinoderms are important both ecologically and geologically. Ecologically, there are few other groupings so abundant in the biotic desert of the deep sea, as well as shallower oceans. Most echinoderms are able to reproduce asexually and regenerate tissue, organs, and limbs; in some cases, they can undergo complete regeneration from a single limb. ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Burgess Shale
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. The rock unit is a black shale and crops out at a number of localities near the town of Field in Yoho National Park Yoho National Park ( ) is a National Parks of Canada, national park of Canada. It is located within the Canadian Rockies, Rocky Mountains along the western slope of the Continental Divide of the Americas in southeastern British Columbia, bordered ... and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42 km to the south. History and significance The Burgess Shale was discovered by palaeontologist Charles Doolittle Walcott, Charles Walcott on 30 August 1909, towards the end of the season's fieldwork. He returned in 1910 with his sons, daughter, and wif ...
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Phyllopod Bed
The Phyllopod bed, designated by USNM locality number 35k, is the most famous fossil-bearing member of the Burgess Shale fossil ''Lagerstätte''. It was quarried by Charles Walcott from 1911–1917 (and later named Walcott Quarry), and was the source of 95% of the fossils he collected during this time; tens of thousands of soft-bodied fossils representing over 150 genera have been recovered from the Phyllopod bed alone. Stratigraphy and location The phyllopod bed is a 2.31 m thick layer of the 7 m thick Greater Phyllopod Bed, found in the Walcott Quarry on Fossil Ridge, between Wapta Mountain and Mount Field, at an elevation of around , around north of the railway town of Field, British Columbia, in the Canadian Rocky Mountains. It is adjacent to Mount Burgess, where Walcott first discovered the Burgess Shale formation. Walcott divided the bed into twelve units based on the rock type and fossil content. Certain fossil beds provide reference levels and can b ...
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Arkarua
''Arkarua adami'' is a small, Precambrian disk-like fossil with a raised center, a number of radial ridges on the rim, and a five-pointed central depression marked with radial lines of five small dots from the middle of the disk center. Fossils range from 3 to 10 mm in diameter. ''Arkarua'' is known only from the Ediacaran beds of the Flinders Ranges in South Australia. The generic name refers to Arkaroo, a giant snake from the Dreaming of the local Aboriginal people. Classification All known specimens of ''Arkarua'' are casts that give no clue to the internal structure, making classification problematic. Because of ''Arkaruas pentamerous symmetry, it is tentatively placed within phylum Echinodermata. Because of its flattened disk- or button-shape, coupled with its pentamerous symmetry, some claim that it can be further classified into the Edrioasteroidea, a class of the echinoderms. This identification remains suspect, as the fossils do not appear to have either madrep ...
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Precambrian
The Precambrian (or Pre-Cambrian, sometimes abbreviated pꞒ, or Cryptozoic) is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of the Phanerozoic Eon, which is named after Cambria, the Latinised name for Wales, where rocks from this age were first studied. The Precambrian accounts for 88% of the Earth's geologic time. The Precambrian is an informal unit of geologic time, subdivided into three eons ( Hadean, Archean, Proterozoic) of the geologic time scale. It spans from the formation of Earth about 4.6 billion years ago ( Ga) to the beginning of the Cambrian Period, about million years ago ( Ma), when hard-shelled creatures first appeared in abundance. Overview Relatively little is known about the Precambrian, despite it making up roughly seven-eighths of the Earth's history, and what is known has largely been discovered from the 1960s onwards. The Precambrian fossil ...
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Stereom
Stereom is a calcium carbonate material that makes up the internal skeletons found in all echinoderms, both living and fossilized forms. It is a sponge-like porous structure which, in a sea urchin may be 50% by volume living cells, and the rest being a matrix of calcite crystals. The size of openings in stereom varies in different species and in different places within the same organism. When an echinoderm becomes a fossil, microscopic examination is used to reveal the structure and such examination is often an important tool to classify the fossil as an echinoderm or related creature. Evolution Stereom was the first form of biomineralization to evolve in deuterostomes, predating the evolution of spicules in tunicates and bone in vertebrates. Stereom likely evolved before other distinctive traits of echinoderms, such as radial symmetry, as it is present in basal echinoderms with bilaterally symmetric or asymmetric body plans. Stereom may have evolved alongside a transition in ocea ...
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Burgess Shale Fossils
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's book '' Wonderful Life'' describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested. The fossil beds are in a series of shale layers, averaging and totalling about in thickness. These layers were deposited against the face of a high undersea limestone cliff. All these features were later raised up above current sea level during the cr ...
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