Vampyropoda
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Vampyropoda
Octopodiformes is a superorder of the subclass Coleoidea, comprising the octopuses and the vampire squid. All living members of Octopodiformes have eight arms, either lacking the two tentacles of squid (as is the case in octopuses) or modifying the tentacles into thin filaments (as in vampire squid). Octopodiformes is often considered the crown group of octopuses and vampire squids, including all descendants of their common ancestor. Some authors use the term Vampyropoda for the same general category, though others use "Vampyropoda" to refer to the total group (all cephalopods closer to octopods than to true squid). Another term is Octobranchia, referring to cephalopods without prominent tentacles. '' Pohlsepia'', originally described as earliest octopod is considered as dubious for this group in later study. The oldest unambiguous vampyropod is ''Syllipsimopodi'', a squid-like cephalopod from the Mississippian-age Bear Gulch Lagerstätte of Montana. ''Syllipsimopodi'' has a com ...
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Octopus
An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefish, and nautiloids. Like other cephalopods, an octopus is bilaterally symmetric with two eyes and a beaked mouth at the center point of the eight limbs. The soft body can radically alter its shape, enabling octopuses to squeeze through small gaps. They trail their eight appendages behind them as they swim. The siphon is used both for respiration and for locomotion, by expelling a jet of water. Octopuses have a complex nervous system and excellent sight, and are among the most intelligent and behaviourally diverse of all invertebrates. Octopuses inhabit various regions of the ocean, including coral reefs, pelagic waters, and the seabed; some live in the intertidal zone and others at abyssal depths. Most species grow quickly, mature ea ...
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Proteroctopus
''Proteroctopus'' is an extinct genus of cephalopod that lived in the Middle Jurassic, approximately 164 million years ago. It is only known from a single species ''P. ribeti''. The single fossil specimen assigned to this species originates from the Lower Callovian of Voulte-sur-Rhône in France. It is currently on display at the Musée de Paléontologie de La Voulte-sur-Rhône. While originally interpreted as an early octopus, a 2016 restudy of the specimen considered it to be a basal member of the Vampyropoda, less closely related to octopus or vampire squid than either of the two groups are to each other. A 2022 phylogenetic analysis found it to be more closely related to vampire squid than to octopuses.om the Mississippian Bear Gulch Lagerstätte sheds light on early vampyropod evolution. A phylogenetic analysis by Kruta et. al indicates that Proteroctopus may be more closely related to the Vampyrpmorpha based on its unique morphology: two fins, head fused to the body, ei ...
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Cephalopoda
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been identified. Tw ...
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Bear Gulch Lagerstätte
The Bear Gulch Limestone is a limestone-rich geological lens in central Montana, renowned for the quality of its late Mississippian-aged fossils. It is exposed over a number of outcrops northeast of the Big Snowy Mountains, and is often considered a component of the more widespread Heath Formation. The Bear Gulch Limestone reconstructs a diverse, though isolated, marine ecosystem which developed near the end of the Serpukhovian age. It is a lagerstätte, a particular type of rock unit with exceptional fossil preservation of both articulated skeletons and soft tissues. Bear Gulch fossils include a variety of fish, invertebrates, and algae occupying a number of different habitats within a preserved shallow bay. Fish include a high diversity of unusual chondrichthyans (cartilaginous fish) and one of the oldest known lampreys, along with other vertebrates. Invertebrates include numerous fossils of crustaceans, worms, cephalopods, and sea sponges, which are concentrated in different p ...
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Syllipsimopodi
''Syllipsimopodi'' is an extinct member of the cephalopod subclass Coleoidea belonging to the clade Vampyropoda, which includes octopuses (Octopoda) and vampire squids (Vampyromorphida). The type and only known species is ''Syllipsimopodi bideni'', named in honor of US President Joe Biden, and to raise awareness of his climate change policies. The well-preserved holotype fossil was found in the Bear Gulch Limestone deposit in the US state of Montana in 1988, and donated that year to the Royal Ontario Museum by one B. Hawes, designated ROMIP 64897. The species lived during the (Carboniferous) mississippian 330.3 to 323.4 million years ago pushing back the group of cephalopods by 81.9 million years. The cephalopod was named after Joe Biden, who was president of the United States at the time of its discovery. Whalen and Landman said in an interview that this name was chosen because they were, “encouraged by his plans to address climate change and to fund scientific research†...
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Stem-group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown gr ...
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Vampyromorphida
Vampyromorphida is an order of cephalopods comprising one known extant species (''Vampyroteuthis infernalis'') and many extinct taxa. Physically, they somewhat resemble octopuses (their closest relatives), but the eight arms are united by a web of skin, and two smaller cilia are also present. Properly speaking, the vampire squid does not possess cilia, but cirri (cilia-like projections). Classification *Order Vampyromorphida **Suborder † Kelaenina ***Family †Muensterellidae **Suborder † Prototeuthina ***Family † Loligosepiidae ***Family † Geopeltididae ***Family † Lioteuthididae ***Family † Mastigophoridae **Suborder † Mesoteuthina ***Family † Palaeololiginidae ****Subfamily † Teudopseinae ****Subfamily † Palaeololigininae **Suborder Vampyromorphina ***Family Vampyroteuthidae The following taxa were long considered to belong to Vampyromorphida, but this placement may be incorrect:Fischer, Jean-Claude & Riou, Bernard (2002): ''Vampyronassa rhodanica'' ...
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Incertae Sedis
' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty at specific taxonomic levels is indicated by ' (of uncertain family), ' (of uncertain suborder), ' (of uncertain order) and similar terms. Examples *The fossil plant '' Paradinandra suecica'' could not be assigned to any family, but was placed ''incertae sedis'' within the order Ericales when described in 2001. * The fossil ''Gluteus minimus'', described in 1975, could not be assigned to any known animal phylum. The genus is therefore ''incertae sedis'' within the kingdom Animalia. * While it was unclear to which order the New World vultures (family Cathartidae) should be assigned, they were placed in Aves ''incertae sedis''. It was later agreed to place them in a separate order, Cathartiformes. * Bocage's longbill, ''Motacilla bocagii' ...
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Muensterellidae
Muensterellidae is a family of stem-octopod cephalopods from the Late Jurassic to Late Cretaceous. Phylogeny Muensterellidae is one of two families in the superfamily Muensterelloidea along with the Patelloctopodidae. The muensterelloids are characterized by having a roughly spoon-shaped end of the gladius ''Gladius'' () is a Latin word meaning "sword" (of any type), but in its narrow sense it refers to the sword of ancient Roman foot soldiers. Early ancient Roman swords were similar to those of the Greeks, called '' xiphe'' (plural; singular ''xi ... called the patella. This type of gladius is likely ancestral to the gladius remnants of modern octopuses. References {{Taxonbar, from=Q30434538 Octopuses Prehistoric cephalopod families ...
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Muensterelloidea
Muensterelloidea is a superfamily (or clade) of stem-octopod cephalopods from the Early Jurassic to Late Cretaceous. Two families are currently identified, Muensterellidae, and Patelloctopodidae. The clade is the ancestral group from which modern octopus arose. Description Overall Soft tissue anatomy within the Muensterelloidea is well-documented. Muenster (1842), who first recorded the holotype specimen of ''Muensterella'', noted its egg-shaped anatomy and apparent lack of swimming fins.Fuchs, D., Keupp, H., & Engeser, T. (2003). New records of soft parts of Muensterella scutellaris Muenster, 1842 (Coleoidea) from the Late Jurassic Plattenkalks of Eichstätt and their significance for octobrachian relationships. Berliner Paläobiologische Abhandlungen, 3, 101-111. Indeed, in all preserved specimens the same body structure is present, with all lacking a distinct swimming fin. Fuchs ''et al.'' (2003) suggested it possessed marginal fins, as a s ...
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Squid
True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting these criteria. Like all other cephalopods, squid have a distinct head, bilateral symmetry, and a mantle. They are mainly soft-bodied, like octopuses, but have a small internal skeleton in the form of a rod-like gladius (cephalopod), gladius or pen, made of chitin. Squid diverged from other cephalopods during the Jurassic and occupy a similar role to teleost fish as open water predators of similar size and behaviour. They play an important role in the open water food web. The two long tentacles are used to grab prey and the eight arms to hold and control it. The beak then cuts the food into suitable size chunks for swallowing. Squid are rapid swimmers, moving by Aquatic locomotion#Jet propulsion, jet propulsion, and largely locate their ...
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