Usnea Filipendula
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Usnea Filipendula
''Usnea filipendula'', the fishbone beard lichen, is a pale gray-green fruticose lichen with a pendant growth form, growing in up to 20 cm many-branching tassels hanging from the bark of trees. In California, it mostly grows on mostly conifer in the Coast Range, but also in the western slopes of the Sierra Nevada range.Field Guide to California Lichens, Stephen Sharnoff, Yale University Press, 2014, It lacks apothecia. It is similar to '' Usnea scabrata'', but is darker, has a thicker cortex, and different chemistry. Lichen spot tests are K+ red, KC−, C−, and P+ yellow. References filipendula ''Filipendula'' is a genus of 12 species of perennial herbaceous flowering plants in the family Rosaceae, native to the temperate regions of the Northern Hemisphere. Well-known species include meadowsweet (''Filipendula ulmaria'') and dropwort ( ... Lichen species Lichens described in 1881 Lichens of Europe Lichens of North America Taxa named by James Stirton {{P ...
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Usnea Scabrata
''Usnea scabrata'', the straw beard lichen, is a pale grayish-yellowish green, slender, pendant, branching from the base, unequally branching, shrubby fruticose lichen that grows from holdfasts on trees.Field Guide to California Lichens, Stephen Sharnoff, Yale University Press, 2014, It is warty with abundant isidia. It resembles ''Usnea filipendula ''Usnea filipendula'', the fishbone beard lichen, is a pale gray-green fruticose lichen with a pendant growth form, growing in up to 20 cm many-branching tassels hanging from the bark of trees. In California, it mostly grows on mostly conife ...''. References scabrata Lichen species Lichens of North America Lichens described in 1875 Taxa named by William Nylander (botanist) {{Parmeliaceae-stub ...
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Lichens Of Europe
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Lichens Described In 1881
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Lichen Species
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Usnea
''Usnea'' is a genus of mostly pale grayish-green fruticose lichens that grow like leafless mini-shrubs or tassels anchored on bark or twigs.Field Guide to California Lichens, Stephen Sharnoff, Yale University Press, 2014, The genus is in the family Parmeliaceae. It grows all over the world. Members of the genus are commonly called old man's beard, beard lichen, or beard moss. Like other lichens it is a symbiosis of two or three fungi and an alga. In ''Usnea'', the fungus belongs to the division Ascomycota, while the alga is a member of the division Chlorophyta. Members of the genus are similar to those of the genus '' Alectoria''. A distinguishing test is that the branches of ''Usnea'' are somewhat elastic, but the branches of ''Alectoria'' snap cleanly off. Systematics The genus ''Usnea'' was circumscribed by Michel Adanson in 1763. He used the name designated by Johann Jacob Dillenius, whose earlier published description did not met the rules of valid publication as estab ...
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Lichen Spot Test
A spot test in lichenology is a spot analysis used to help identify lichens. It is performed by placing a drop of a chemical on different parts of the lichen and noting the colour change (or lack thereof) associated with application of the chemical. The tests are routinely encountered in dichotomous keys for lichen species, and they take advantage of the wide array of lichen products produced by lichens and their uniqueness among taxa. As such, spot tests reveal the presence or absence of chemicals in various parts of a lichen. They were first proposed by the botanist William Nylander in 1866. Three common spot tests use either 10% aqueous KOH solution (K test), saturated aqueous solution of bleaching powder or calcium hypochlorite (C test), or 5% alcoholic ''p''-phenylenediamine solution (P test). The colour changes occur due to presence of particular secondary metabolites in the lichen. There are several other less frequently used spot tests of more limited use that are employed ...
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Cortex (lichen)
In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. The cortex is composed mostly of large thin-walled parenchyma cells of the ground tissue system and shows little to no structural differentiation. The outer cortical cells often acquire irregularly thickened cell walls, and are called collenchyma cells. Plants Stems and branches In the three dimensional structure of herbaceous stems, the epidermis, cortex and vascular cambium form concentric cylinders around the inner cylindrical core of pith. Some of the outer cortical cells may contain chloroplasts, giving them a green color. They can therefore produce simple carbohydrates through photosynthesis. In woody plants, the cortex is located between the periderm (bark) and the vascular tissue (phloem, in particular). It is responsible for the transportation of materials into the central cylinder of the root through diffusion and may also be ...
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Apothecia
An ascocarp, or ascoma (), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are most commonly bowl-shaped (apothecia) but may take on a spherical or flask-like form that has a pore opening to release spores (perithecia) or no opening (cleistothecia). Classification The ascocarp is classified according to its placement (in ways not fundamental to the basic taxonomy). It is called ''epigeous'' if it grows above ground, as with the morels, while underground ascocarps, such as truffles, are termed ''hypogeous''. The structure enclosing the hymenium is divided into the types described below (apothecium, cleistothecium, etc.) and this character ''is'' important for the taxonomic classification of the fungus. Apothecia can be relatively large and fleshy, whereas the others are microscopic—about the size of flecks of ...
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Fruticose Lichen
A fruticose lichen is a form of lichen fungi that is characterized by a coral-like shrubby or bushy growth structure. It is formed from a symbiotic relationship of a photobiont such as green algae or less commonly cyanobacteria and one, two or more mycobionts. Fruticose lichens are not a monophyletic and holophyletic lineage, but is a form encountered in many classes. Fruticose lichens have a complex vegetation structure, and are characterized by an ascending, bushy or pendulous appearance. As with other lichens, many fruticose lichens can endure high degrees of desiccation. They grow slowly and often occur in habitats such as on tree barks, on rock surfaces and on soils in the Arctic and mountain regions. Characteristics Fruticose lichens are lichens composed of a shrubby or bushy thallus and a holdfast. The thallus is the vegetative body of a lichen that does not have true leaves, stems, or roots. The thallus colour is affected by the algae in the lichen, compounds created by t ...
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Sierra Nevada
The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily in Nevada. The Sierra Nevada is part of the American Cordillera, an almost continuous chain of mountain ranges that forms the western "backbone" of the Americas. The Sierra runs north-south and its width ranges from to across east–west. Notable features include General Sherman, the largest tree in the world by volume; Lake Tahoe, the largest alpine lake in North America; Mount Whitney at , the highest point in the contiguous United States; and Yosemite Valley sculpted by glaciers from one-hundred-million-year-old granite, containing high waterfalls. The Sierra is home to three national parks, twenty wilderness areas, and two national monuments. These areas include Yosemite, Sequoia, and Kings Canyon National Parks; and Devils Po ...
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Coast Range (EPA Ecoregion)
The Coast Range ecoregion is a Level III ecoregion designated by the Environmental Protection Agency (EPA) in the U.S. states of Washington, Oregon, and California. It stretches along the Pacific Coast from the tip of the Olympic Peninsula in the north to the San Francisco Bay in the south, including Grays Harbor, Willapa Bay, and the Long Beach Peninsula in Washington, the entire length of the Oregon Coast, and the Northern California Coast. Named for the Coast Range mountains, it encompasses the lower elevations of the Olympic Mountains, the Oregon Coast Range, the Californian North Coast Ranges, and surrounding lowlands. The low mountains of the ecoregion are covered by highly productive, rain-drenched evergreen forests that are home to the three tallest conifer species in the world: Coast redwood, Coast Douglas-fir, and Sitka spruce. Historically, Sitka spruce forests dominated the fog-shrouded coast, while a mosaic of western redcedar, western hemlock, and seral Dou ...
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