Uranocentrodon
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Uranocentrodon
''Uranocentrodon'' is an extinct genus of the family Rhinesuchidae. Known from a skull, ''Uranocentrodon'' was a large predator with a length up to . Originally named ''Myriodon'' by van Hoepen in 1911, it was transferred to a new genus on account of the name being preoccupied in 1917. It has been synonymized with ''Rhinesuchus'', but this has not been widely supported. It was also originally considered to be of Triassic age, but more recent analysis has placed its age as just below the Permian-Triassic boundary. History and specimens Fossils of the animal now known as ''Uranocentrodon'' were first discovered in a sandstone quarry in the Normandien Formation near Senekal in Free State, South Africa. The first example of these fossils was a fragmentary skull excavated by a local family. Subsequent excavations and searches undertaken by various parties unearthed more specimens which hailed from the quarry. Among these specimens included three nearly complete skeletons overlappi ...
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Rhinesuchidae
Rhinesuchidae is a family of tetrapods that lived primarily in the Permian period. They belonged to the broad group Temnospondyli, a successful and diverse collection of semiaquatic tetrapods which modern amphibians are probably descended from. Rhinesuchids can be differentiated from other temnospondyls by details of their skulls, most notably the interior structure of their otic notches at the back of the skull. They were among the earliest-diverging members of the Stereospondyli, a subgroup of temnospondyls with flat heads and aquatic habits. Although more advanced stereospondyls evolved to reach worldwide distribution in the Triassic period, rhinesuchids primarily lived in the high-latitude environments of Gondwana (what is now South America and Africa) during the Guadalupian and Lopingian epochs of the Permian. The taxonomy of this family has been convoluted, with more than twenty species having been named in the past; a 2017 review recognized only eight of them (distributed a ...
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Changhsingian
In the geologic time scale, the Changhsingian or Changxingian is the latest age or uppermost stage of the Permian. It is also the upper or latest of two subdivisions of the Lopingian Epoch or Series. The Changhsingian lasted from to 251.902 million years ago (Ma). It was preceded by the Wuchiapingian and followed by the Induan. The greatest mass extinction in the Phanerozoic eon, the Permian–Triassic extinction event, occurred during this age. The extinction rate peaked about a million years before the end of this stage. Stratigraphic definitions The Changhsingian is named after Changxing () in northern Zhejiang, China. The stage was named for the Changhsing Limestone. The name was first used for a stage in 1970; 1973: ''Permian stages names'', in: : ''The Permian and Triassic systems and their mutual boundary'', Canadian Society of Petroleum Geologists Memoir 2, pp 522–548. and was anchored in the international timescale in 1981.; 2006: ''The Global Boundary Stratotype Sect ...
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National Museum, Bloemfontein
The National Museum in Bloemfontein Bloemfontein, ( ; , "fountain of flowers") also known as Bloem, is one of South Africa's three capital cities and the capital of the Free State province. It serves as the country's judicial capital, along with legislative capital Cape To ... dates back to the 1870s, when its collections and displays comprised mainly rarities from around the world. It was established on 20 July 1877 and opened to the public on 20 May 1878. The old Raadsaal building was made available by the then Free State Volksraad for hosting the Museum collections. A new museum building was completed in 1913 and the official opening took place in 1915. Since 1881 the museum has been subsidised mainly by government. Through donations, purchases and exchange agreements the Museum’s collections slowly got established and grew to a wide variety of natural and human science artefacts. The first honorary curator was Dr Hugh Exton. The first director, Dr E.C.N. van Hoepe ...
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Lungfish
Lungfish are freshwater vertebrates belonging to the order Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, including the presence of lobed fins with a well-developed internal skeleton. Lungfish represent the closest living relatives of the tetrapods. Today there are only six known species of lungfish, living in Africa, South America, and Australia. The fossil record shows that lungfish were abundant since the Triassic. While vicariance would suggest this represents an ancient distribution limited to the Mesozoic supercontinent Gondwana, the fossil record suggests advanced lungfish had a widespread freshwater distribution and the current distribution of modern lungfish species reflects extinction of many lineages subsequent to the breakup of Pangaea, Gondwana and Laurasia. Lungfish have historically been referred to as salamanderfish, but this t ...
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Branchiosauridae
Branchiosauridae is an extinct family of small amphibamiform temnospondyls with external gills and an overall juvenile appearance. The family has been characterized by hundreds of well-preserved specimens from the Permo-Carboniferous of Middle Europe.Schoch, R.R. 2008. The intrarelationships and evolutionary history of the temnospondyl family branchiosauridae. Journal of Systematic Palaeontology. 6(4):409-431. Specimens represent well defined ontogenetic stages and thus the taxon has been described to display paedomorphy (perennibranchiate). However, more recent work has revealed branchiosaurid taxa that display metamorphosing trajectories.Schoch, R.R. and Frobisch, N.B. 2006. Alternative Pathways in an Extinct Amphibian Clade. Evolution. 60(7):1467-1475 The name Branchiosauridae (“Branchio” in Ancient Greek denoting gills and “saurus” meaning lizard) refers to the retention of gills. Geological/paleoenvironmental information Branchiosaurids mostly inhabited Permo-Carbo ...
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Dvinosaurus
''Dvinosaurus'' is an extinct genus of amphibious temnospondyls localized to regions of western and central Russia during the middle and late Permian, approximately 265-254 million years ago. Its discovery was first noted in 1921 by Russian paleontologist Vladimir Prokhorovich Amalitskii in a posthumously published paper that documents the findings of a site in Russia's Arkhangelsk District. Its name is derived from the proximity of this site to the Northern Dvina River. ''Dvinosaurus'' is thought to have been a gill-breathing, fully-aquatic tetrapod, characterized by a large, triangular head, short limbs, and a long powerful tail. A typical individual could grow to be approximately 40 in (100 cm) in length. Within this genus, the number of documented species has varied over the years since its discovery. Prior to his untimely death, Amalitskii described three species, ''Dvinosaurus primus, Dvinosaurus secundus,'' and ''Dvinosaurus tertius.'' Upon further analysis, however, ...
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Dvinosauria
Dvinosaurs are one of several new clades of Temnospondyl amphibians named in the phylogenetic review of the group by Yates and Warren 2000. They represent a group of primitive semi-aquatic to completely aquatic amphibians, and are known from the Late Carboniferous to the Early Triassic, being most common in the Permian period. Their distinguishing characteristics are a reduction of the otic notch; the loss of a flange on the rear side of the pterygoid; and 28 or more presacral vertebrae. Trimerorhachidae is the most basal family of dvinosaurs. Most other dvinosaurs are placed in the superfamily Dvinosauroidea. Within Dvinosauroidea are two families, Eobrachyopidae and Tupilakosauridae, as well as dvinosaurs that cannot be placed in either family, such as '' Dvinosaurus'' and ''Kourerpeton''. A 2008 phylogenetic analysis found Eobrachyopidae to be paraphyletic, representing a grade of basal dvinosauroids. Below is a cladogram showing the phylogenetic relationships of dvinos ...
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Axolotl
The axolotl (; from nci, āxōlōtl ), ''Ambystoma mexicanum'', is a paedomorphic salamander closely related to the tiger salamander. Axolotls are unusual among amphibians in that they reach adulthood without undergoing metamorphosis. Instead of taking to the land, adults remain aquatic and gilled. The species was originally found in several lakes underlying what is now Mexico City, such as Lake Xochimilco and Lake Chalco. These lakes were drained by Spanish settlers after the conquest of the Aztec Empire, leading to the destruction of much of the axolotl’s natural habitat. Axolotls should not be confused with the larval stage of the closely related tiger salamander (''A. tigrinum''), which are widespread in much of North America and occasionally become paedomorphic. Neither should they be confused with mudpuppies (''Necturus'' spp.), fully aquatic salamanders from a different family that are not closely related to the axolotl but bear a superficial resemblance. , wild ax ...
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Necturus
''Necturus'' is a genus of aquatic salamanders native to the eastern United States and Canada. They are commonly known as waterdogs and mudpuppies. The common mudpuppy ''(N. maculosus)'' is probably the best-known species – as an amphibian with gill slits, it is often dissected in comparative anatomy classes. Taxonomy The genus is under scrutiny by herpetologists. The relationship between the species is still being studied. In 1991 Collins elevated ''N. maculosus louisianensis'' to full species status, usually considered a subspecies of the common mudpuppy (''N. maculosus''), but his interpretation was not largely followed.Petranka, J.W. (1998). Salamanders of the United States and Canada. Smithsonian Institution Press . However, a 2018 study confirmed it as a distinct species, with Amphibian Species of the World following these results, although other authorities do not.'''' Species There are seven to eight species: Two known fossil species, '' N. krausei'' and an unna ...
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Neoteny
Neoteny (), also called juvenilization,Montagu, A. (1989). Growing Young. Bergin & Garvey: CT. is the delaying or slowing of the physiological, or somatic, development of an organism, typically an animal. Neoteny is found in modern humans compared to other primates. In progenesis or paedogenesis, sexual development is accelerated. Both neoteny and progenesis result in paedomorphism (as having the form typical of children) or paedomorphosis (changing towards forms typical of children), a type of heterochrony. It is the retention in adults of traits previously seen only in the young. Such retention is important in evolutionary biology, domestication and evolutionary developmental biology. Some authors define paedomorphism as the retention of larval traits, as seen in salamanders.Schell, S. C. ''Handbook of Trematodes of North America North of Mexico'', 1985, pg. 22 History and etymology The origins of the concept of neoteny have been traced to the Bible (as argued by Ashley Monta ...
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Salamander
Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela. Salamander diversity is highest in eastern North America, especially in the Appalachian Mountains; most species are found in the Holarctic realm, with some species present in the Neotropical realm. Salamanders rarely have more than four toes on their front legs and five on their rear legs, but some species have fewer digits and others lack hind limbs. Their permeable skin usually makes them reliant on habitats in or near water or other cool, damp places. Some salamander species are fully aquatic throughout their lives, some take to the water intermittently, and others are entirely terrestrial as adults. This group of amphibians is capable of regenerating lost lim ...
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External Gills
External gills are the gills of an animal, most typically an amphibian, that are exposed to the environment, rather than set inside the pharynx and covered by gill slits, as they are in most fishes. Instead, the respiratory organs are set on a frill of stalks protruding from the sides of an animal's head. This type of gill is most commonly observed on the aquatic larva of most species of salamanders, lungfish, and bichirs (which have only one large pair), and are retained by neotenic adult salamanders and some species of adult lungfish. They are present on non-transforming salamander species, such as most members of the family Proteidae (the olm and mudpuppies) and the family Sirenidae, which naturally never metamorphose into an air-breathing form. The embryos of frogs and caecilians also develop external gills at some point in their development, though these are either resorbed before or disappear shortly after hatching. Fossils of the distant relatives of modern amphibians, such ...
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