Triassurus
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Triassurus
''Triassurus'' is an extinct genus of amphibian, and the oldest member of Caudata (salamanders and close relalatives). It is known from the Middle to Upper Triassic (Ladinian-Carnian) aged Madygen Formation in Kyrgyzstan. The type species is ''T. sixtelae''. Description The holotype specimen is a partial and poorly preserved skeleton (PIN-2584/10), including the skull. Holotype specimen had tiny skull just long, but it was probably a larva: the neural arches of the vertebrae were still paired and no vertebral centers show any degree of ossification. The skull closely resembles that of current salamanders, especially as regards the long space in the joint of the maxillary bones and the lack of bone connection between the pterygoid and the maxilla. There were about 20 presacral vertebrae, while the legs were small. In 2020, a more complete adult specimen (FG 596/V/20) with long skull was described. Classification ''Triassurus'' was first described in 1978, based on a fossil foun ...
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Madygen Formation
The Madygen Formation (Russian: Madygen Svita) is a Late Triassic (Carnian) geologic formation and Lagerstätte in the Batken and Osh Regions of western Kyrgyzstan, with minor outcrops in neighboring Tajikistan and Uzbekistan. The conglomerates, sandstones and mudstones of the thick formation were deposited in terrestrial lacustrine, alluvial, fluvial and deltaic environments. The formation, extending across the Fergana Valley and Fergana Range, is unique for Central Asia, as it represents one of the few known continental deposits and the Madygen Formation is renowned for the preservation of more than 20,000 fossil insects, making it one of the richest Triassic Lagerstätten in the world. Other vertebrate fossils as fish, amphibians, reptiles and synapsids have been recovered from the formation too, as well as minor fossil flora. The lake sediments of the Lagerstätte provided fossil cartilaginous fishes and their egg capsules and unusual Triassic reptiles like ''Sharovipteryx'' ...
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Caudata
The Caudata are a group of amphibians containing the extant salamanders (Urodela) and all extinct species of amphibians more closely related to salamanders than to frogs. They are typically characterized by a superficially lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. Disagreement exists between different authorities as to the definition of the terms "Caudata" and "Urodela". Some maintain that Urodela should be restricted to the crown group, with Caudata being used for the total group. Others restrict the name Caudata to the crown group and use Urodela for the total group. The former approach seems to be most widely adopted and is used in this article. Evolution The origins and evolutionary relationships between the three main groups of amphibians ( apodans, urodeles and anurans) is a matter of debate. A 2005 molecular phylogeny, based on rDNA analysis, suggest ...
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Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', t ...
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Temnospondyl
Temnospondyli (from Greek language, Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order (biology), order of small to giant tetrapods—often considered Labyrinthodontia, primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had cha ...
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Fossils Of Kyrgyzstan
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolute ...
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Scapula
The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on either side of the body being roughly a mirror image of the other. The name derives from the Classical Latin word for trowel or small shovel, which it was thought to resemble. In compound terms, the prefix omo- is used for the shoulder blade in medical terminology. This prefix is derived from ὦμος (ōmos), the Ancient Greek word for shoulder, and is cognate with the Latin , which in Latin signifies either the shoulder or the upper arm bone. The scapula forms the back of the shoulder girdle. In humans, it is a flat bone, roughly triangular in shape, placed on a posterolateral aspect of the thoracic cage. Structure The scapula is a thick, flat bone lying on the thoracic wall that provides an attachment for three groups of muscles: intrin ...
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Squamosal Bone
The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestral component of the dermal roof and is typically thin compared to other skull bones. The squamosal bone lies ventral to the temporal series and otic notch, and is bordered anteriorly by the postorbital. Posteriorly, the squamosal articulates with the quadrate and pterygoid bones. The squamosal is bordered anteroventrally by the jugal and ventrally by the quadratojugal. Function in reptiles In reptiles, the quadrate and articular bones of the skull articulate to form the jaw joint. The squamosal bone lies anterior to the quadrate bone. Anatomy in synapsids Non-mammalian synapsids In non-mammalian synapsids, the jaw is composed of four bony elements and referred to as a quadro-articular jaw because the joint is between the articular an ...
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Parietal Bone
The parietal bones () are two bones in the Human skull, skull which, when joined at a fibrous joint, form the sides and roof of the Human skull, cranium. In humans, each bone is roughly quadrilateral in form, and has two surfaces, four borders, and four angles. It is named from the Latin ''paries'' (''-ietis''), wall. Surfaces External The external surface [Fig. 1] is convex, smooth, and marked near the center by an eminence, the parietal eminence (''tuber parietale''), which indicates the point where ossification commenced. Crossing the middle of the bone in an arched direction are two curved lines, the superior and inferior temporal lines; the former gives attachment to the temporal fascia, and the latter indicates the upper limit of the muscular origin of the temporal muscle. Above these lines the bone is covered by a tough layer of fibrous tissue – the epicranial aponeurosis; below them it forms part of the temporal fossa, and affords attachment to the temporal muscle. ...
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Parasphenoid
The parasphenoid is a bone which can be found in the cranium of many vertebrates. It is an unpaired dermal bone which lies at the midline of the roof of the mouth. In many reptiles (including birds), it fuses to the endochondral (cartilage-derived) basisphenoid bone of the lower braincase, forming a bone known as the parabasisphenoid. Early mammals have a small parasphenoid, but for the most part its function has been replaced by the vomer bone. The parasphenoid has been lost in placental mammals and caecilian amphibians. See also *Ossification of frontal bone *Terms for anatomical location Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ... References Bones of the head and neck {{musculoskeletal-stub ...
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Synapomorphy And Apomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis Th ...
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Branchiosaur
Branchiosauridae is an extinct family of small amphibamiform temnospondyls with external gills and an overall juvenile appearance. The family has been characterized by hundreds of well-preserved specimens from the Permo-Carboniferous of Middle Europe.Schoch, R.R. 2008. The intrarelationships and evolutionary history of the temnospondyl family branchiosauridae. Journal of Systematic Palaeontology. 6(4):409-431. Specimens represent well defined ontogenetic stages and thus the taxon has been described to display paedomorphy (perennibranchiate). However, more recent work has revealed branchiosaurid taxa that display metamorphosing trajectories.Schoch, R.R. and Frobisch, N.B. 2006. Alternative Pathways in an Extinct Amphibian Clade. Evolution. 60(7):1467-1475 The name Branchiosauridae (“Branchio” in Ancient Greek denoting gills and “saurus” meaning lizard) refers to the retention of gills. Geological/paleoenvironmental information Branchiosaurids mostly inhabited Permo-Carbo ...
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Longisquama
''Longisquama'' is a genus of extinct reptile. There is only one species, ''Longisquama insignis'', known from a poorly preserved skeleton and several incomplete fossil impressions from the Middle to Late Triassic Madygen Formation in Kyrgyzstan. It is known from the type fossil specimen, slab and counterslab (PIN 2548/4 and PIN 2584/5) and five referred specimens of possible integumentary appendages (PIN 2584/7 through 9). All specimens are in the collection of the Paleontological Institute of the Russian Academy of Sciences in Moscow. ''Longisquama'' means "long scales"; the specific name ''insignis'' refers to its small size. The ''Longisquama'' holotype is notable for a number of long structures that appear to grow from its skin. The current opinion is that ''Longisquama'' is an ambiguous diapsid and has no bearing on the origin of birds. History Interpretation Researchers Haubold and Buffetaut believed that the structures were long, modified scales attached in pairs t ...
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