Thousand Cankers Disease
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Thousand Cankers Disease
Thousand cankers disease (TCD) is a recently recognized disease of certain walnuts (''Juglans'' spp.). The disease results from the combined activity of the walnut twig beetle (''Pityophthorus juglandis'') and a canker producing fungus, '' Geosmithia morbida''. Until July 2010 the disease was only known to the western United States where over the past decade it has been involved in several large scale die-offs of walnut, particularly black walnut, ''Juglans nigra''. However, in late July 2010 a well-established outbreak of the disease was found in the Knoxville, Tennessee area. This new finding is the first locating it within the native range of its susceptible host, black walnut. In 2013, an outbreak was found in the Veneto region of Italy, where the disease has been found on both black walnut and English walnut. Description Adult walnut twig beetles carry spores of the ''Geosmithia morbida'' fungus, which grows profusely around the pupal chamber of the beetles. Following emerg ...
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Geosmithia Morbida
''Geosmithia morbida'' is a species of anamorphic fungus in the Bionectriaceae family that, together with the activity of the walnut twig beetle, causes thousand cankers disease in species of walnut trees (''Juglans spp.''). It was described as new to science in 2010 from specimens collected in the southern United States. The fungus, transmitted by the walnut twig beetle, ''Pityophthorus juglandis'', is known from the western USA from California to Colorado. The cankers resulting from infection restrict nutrient flow and typically kill the host tree within three to four years. Based on closeness of internal transcribed spacer DNA, the closest relative of ''G. morbida'' is '' G. fassatiae''. The specific epithet ''morbida'' refers to the deadly pathogenic In biology, a pathogen ( el, πάθος, "suffering", "passion" and , "producer of") in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an inf ...
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Phloem
Phloem (, ) is the living biological tissue, tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This transport process is called translocation. In trees, the phloem is the innermost layer of the bark (botany), bark, hence the name, derived from the Ancient Greek word (''phloiós''), meaning "bark". The term was introduced by Carl Nägeli in 1858. Structure Phloem tissue consists of conducting Cell (biology), cells, generally called sieve elements, Ground tissue#Parenchyma, parenchyma cells, including both specialized companion cells or albuminous cells and unspecialized cells and supportive cells, such as fibres and sclereids. Conducting cells (sieve elements) Sieve elements are the type of cell that are responsible for transporting sugars throughout the plant. At maturity they lack a Cell nucleus, nucleus and have very few organelles, ...
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Juglans Microcarpa
''Juglans microcarpa'', known also as the little walnut, Texas walnut, Texas black walnut or little black walnut (as it belongs to the "black walnuts" section ''Juglans'' sect. ''Rhysocaryon''), is a large shrub or small tree (10–30 ft tall) which grows wild along streams and ravines in Texas, New Mexico, Oklahoma, and Kansas, and the northernmost states of Mexico. It produces nuts with a width of 1/2—3/4 in. The pinnately compound leaves bear 7—25 untoothed to finely-toothed leaflets, each 1/4—1/2 in wide. It is found at elevations ranging from 700 ft to 6700 ft. Two varieties are recognized: ''J. microcarpa'' var. ''microcarpa'' and ''J. microcarpa'' var. ''stewartii''. Where the range of ''J. microcarpa'' overlaps with '' J. major'', the two species interbreed, producing populations with intermediate characteristics. This phenomenon has also been found where ''J. microcarpa'' trees grows near '' J. nigra trees ...
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Juglans Californica
''Juglans californica'', the California black walnut, also called the California walnut, or the Southern California black walnut, is a large shrub or small tree (about 20-49 feet tall) of the walnut family, Juglandaceae, endemic to Southern California. Distribution ''Juglans californica'' is generally found in the southern California Coast Ranges, Transverse Ranges, and Peninsular Ranges. It grows as part of mixed woodlands, and also on slopes and in valleys wherever conditions are favorable. It is threatened by development and overgrazing. Some native stands remain in urban Los Angeles in the Santa Monica Mountains and Hollywood Hills. ''J. californica'' grows in riparian woodlands, either in single species stands or mixed with California's oaks (''Quercus'' spp.) and cottonwoods (''Populus fremontii''). Description ''Juglans californica'' can be either a large shrub with 1–5 trunks, or a small, single-trunked tree. The main trunk can fork close to the ground, making it loo ...
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Juglans Major
''Juglans major'' (literally, the larger walnut), also known as Arizona walnut, is a walnut tree which grows to with a DBH of up to at elevations of in Texas, Oklahoma, New Mexico, Arizona, and Utah. It also occurs in Mexico as far south as Guerrero. Common names include Arizona black walnut (as it belongs to the "black walnuts" section ''Juglans'' sect. ''Rhysocaryon''), and the Spanish name ''nogal cimarrón'' (cimarron walnut). Description In moister areas, the tree features a single, stout trunk; there are usually several slender trunks in drier situations. The 8–14 in long pinnately compound leaves bear 9–15 lanceolate leaflets, wide by long. The small nut has a thick shell with deep grooves enclosing an oily, edible seed. Where the range of ''J. major'' overlaps that of '' J. microcarpa'', the two interbreed, producing many intermediate forms.Powell, A. Michael. (1988). ''Trees & shrubs of Trans-Pecos Texas including Big Bend and Guadalupe Mounta ...
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Susceptibility (botany)
Susceptibility may refer to: Physics and engineering In physics the susceptibility is a quantification for the change of an extensive property under variation of an intensive property. The word may refer to: * In physics, the susceptibility of a material or substance describes its response to an applied field. For example: ** Magnetic susceptibility ** Electric susceptibility * The two types of susceptibility above are examples of a linear response function; sometimes the terms ''susceptibility'' and ''linear response function'' are used interchangeably. *In electromagnetic compatibility (EMC), susceptibility is the sensitivity of a device's function to incoming electromagnetic interference Health and medicine * In epidemiology, a susceptible individual is a member of a population who is at risk of becoming infected by a disease * In microbiology, pharmacology, and medicine drug susceptibility is the ability of a microorganism to be inhibited or killed by the drug, as in antibiot ...
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Host (biology)
In biology and medicine, a host is a larger organism that harbours a smaller organism; whether a parasite, parasitic, a mutualism (biology), mutualistic, or a commensalism, commensalist ''guest'' (symbiont). The guest is typically provided with nourishment and shelter. Examples include animals playing host to parasitic worms (e.g. nematodes), cell (biology), cells harbouring pathogenic (disease-causing) viruses, a Fabaceae, bean plant hosting mutualistic (helpful) Rhizobia, nitrogen-fixing bacteria. More specifically in botany, a host plant supplies nutrient, food resources to micropredators, which have an evolutionarily stable strategy, evolutionarily stable relationship with their hosts similar to ectoparasitism. The host range is the collection of hosts that an organism can use as a partner. Symbiosis Symbiosis spans a wide variety of possible relationships between organisms, differing in their permanence and their effects on the two parties. If one of the partners in an ass ...
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Wilting
Wilting is the loss of rigidity of non-woody parts of plants. This occurs when the turgor pressure in non-lignified plant cells falls towards zero, as a result of diminished water in the cells. Wilting also serves to reduce water loss, as it makes the leaves expose less surface area. The rate of loss of water from the plant is greater than the absorption of water in the plant. The process of wilting modifies the leaf angle distribution of the plant (or canopy) towards more erectophile conditions. Lower water availability may result from: * drought conditions, where the soil moisture drops below conditions most favorable for plant functioning; * the temperature falls to the point where the plant's vascular system cannot function; * high salinity, which causes water to diffuse from the plant cells and induce shrinkage; * saturated soil conditions, where roots are unable to obtain sufficient oxygen for cellular respiration, and so are unable to transport water into the plant; or ...
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Symptom
Signs and symptoms are the observed or detectable signs, and experienced symptoms of an illness, injury, or condition. A sign for example may be a higher or lower temperature than normal, raised or lowered blood pressure or an abnormality showing on a medical scan. A symptom is something out of the ordinary that is experienced by an individual such as feeling feverish, a headache or other pain or pains in the body. Signs and symptoms Signs A medical sign is an objective observable indication of a disease, injury, or abnormal physiological state that may be detected during a physical examination, examining the patient history, or diagnostic procedure. These signs are visible or otherwise detectable such as a rash or bruise. Medical signs, along with symptoms, assist in formulating diagnostic hypothesis. Examples of signs include elevated blood pressure, nail clubbing of the fingernails or toenails, staggering gait, and arcus senilis and arcus juvenilis of the eyes. Indicati ...
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Bark Beetle
A bark beetle is the common name for the subfamily of beetles Scolytinae. Previously, this was considered a distinct family (Scolytidae), but is now understood to be a specialized clade of the "true weevil" family (Curculionidae). Although the term "bark beetle" refers to the fact that many species feed in the inner bark (phloem) layer of trees, the subfamily also has many species with other lifestyles, including some that bore into wood, feed in fruit and seeds, or tunnel into herbaceous plants. Well-known species are members of the type genus ''Scolytus'', namely the European elm bark beetle ''S. multistriatus'' and the large elm bark beetle ''S. scolytus'', which like the American elm bark beetle ''Hylurgopinus rufipes'', transmit Dutch elm disease fungi (''Ophiostoma''). The mountain pine beetle ''Dendroctonus ponderosae'', southern pine beetle ''Dendroctonus frontalis'', and their near relatives are major pests of conifer forests in North America. A similarly aggressive spe ...
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Nutrient
A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excreted by cells to create non-cellular structures, such as hair, scales, feathers, or exoskeletons. Some nutrients can be metabolically converted to smaller molecules in the process of releasing energy, such as for carbohydrates, lipids, proteins, and fermentation products (ethanol or vinegar), leading to end-products of water and carbon dioxide. All organisms require water. Essential nutrients for animals are the energy sources, some of the amino acids that are combined to create proteins, a subset of fatty acids, vitamins and certain minerals. Plants require more diverse minerals absorbed through roots, plus carbon dioxide and oxygen absorbed through leaves. Fungi live on dead or living organic matter and meet nutrient needs from their host. ...
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Bark (botany)
Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines, and shrubs. Bark refers to all the tissues outside the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner bark, which in older stems is living tissue, includes the innermost layer of the periderm. The outer bark on older stems includes the dead tissue on the surface of the stems, along with parts of the outermost periderm and all the tissues on the outer side of the periderm. The outer bark on trees which lies external to the living periderm is also called the rhytidome. Products derived from bark include bark shingle siding and wall coverings, spices and other flavorings, tanbark for tannin, resin, latex, medicines, poisons, various hallucinogenic chemicals and cork. Bark has been used to make cloth, canoes, and ropes and used as a surface for paintings and map making. A number of plants a ...
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