Tarachoptera
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Tarachoptera
Tarachoptera is an extinct order of insects, currently solely known from the mid Cretaceous aged Burmese amber. It belongs to Amphiesmenoptera alongside living Lepidopterans (butterflies and moths) and Trichoptera (caddiesflies), but is outside the clade containing the two groups, with an estimated divergence during the Upper Triassic, and therefore over a 100 million year ghost lineage. It currently contains only one family, the Tarachocelidae, which was named in a publication in 2017 a few months before the order itself was published. As the manuscript was in submission, an additional specimen belonging to a new genus justified creating a separate order. Additional species were described in 2018 and 2020. A notable character is the possession of wing scales like members of Lepidoptera, these were initially suggested to have been evolved in parallel evolution from hairs or setae, but a later study suggested that the scales of all amphiesmenopterans are homologous, sharing a commo ...
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Amphiesmenoptera
Amphiesmenoptera is an insect superorder, established by S. G. Kiriakoff, but often credited to Willi Hennig in his revision of insect taxonomy for two sister orders: Lepidoptera (butterflies and moths) and Trichoptera (caddisflies). In 2017, a third fossil order was added to the group, the Tarachoptera. Trichoptera and Lepidoptera share a number of derived characters (synapomorphies) which demonstrate their common descent: * Females, rather than males, are heterogametic (i.e. their sex chromosomes differ). * Dense setae are present in the wings (modified into scales in Lepidoptera). * There is a particular venation pattern on the forewings (the double-looped anal veins). * Larvae have mouth structures and glands to make and manipulate silk. Thus these two extant orders are sisters, with Tarachoptera basal to both groups. Amphiesmenoptera probably evolved in the Jurassic. Lepidoptera differ from the Trichoptera in several features, including wing venation, form of the scales on ...
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Lepidoptera
Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic rank, superfamilies, 10 percent of the total described species of living organisms. It is one of the most widespread and widely recognizable insect orders in the world. The Lepidoptera show many variations of the basic body structure that have evolved to gain advantages in lifestyle and distribution. Recent estimates suggest the order may have more species than earlier thought, and is among the four most wikt:speciose, speciose orders, along with the Hymenoptera, fly, Diptera, and beetle, Coleoptera. Lepidopteran species are characterized by more than three derived features. The most apparent is the presence of scale (anatomy), scales that cover the torso, bodies, wings, and a proboscis. The scales are modified, flattened "hairs", and give ...
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Burmese Amber
Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The amber is of significant palaeontological interest due to the diversity of flora and fauna contained as inclusions, particularly arthropods including insects and arachnids but also birds, lizards, snakes, frogs and fragmentary dinosaur remains. The amber has been known and commercially exploited since the first century AD, and has been known to science since the mid-nineteenth century. Research on the deposit has attracted controversy due to its alleged role in funding internal conflict in Myanmar and hazardous working conditions in the mines where it is collected. Geological context, depositional environment and age The amber is found within the Hukawng Basin, a large Cretaceous-Cenozoic sedimentary basin within northern Myanmar. The s ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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Trichoptera
The caddisflies, or order Trichoptera, are a group of insects with aquatic larvae and terrestrial adults. There are approximately 14,500 described species, most of which can be divided into the suborders Integripalpia and Annulipalpia on the basis of the adult mouthparts. Integripalpian larvae construct a portable casing to protect themselves as they move around looking for food, while Annulipalpian larvae make themselves a fixed retreat in which they remain, waiting for food to come to them. The affinities of the small third suborder Spicipalpia are unclear, and molecular analysis suggests it may not be monophyletic. Also called sedge-flies or rail-flies, the adults are small moth-like insects with two pairs of hairy membranous wings. They are closely related to the Lepidoptera (moths and butterflies) which have scales on their wings; the two orders together form the superorder Amphiesmenoptera. The aquatic larvae are found in a wide variety of habitats such as streams, river ...
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Upper Triassic
The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', the middle Muschelkalk (shell-bearing limestone) and the upper Keuper (coloured clay). The Late Triassic Series co ...
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Ghost Lineage
A ghost lineage is a hypothesized ancestor in a species lineage that has left no fossil evidence yet can be inferred to exist because of gaps in the fossil record or genomic evidence. The process of determining a ghost lineage relies on fossilized evidence before and after the hypothetical existence of the lineage and extrapolating relationships between organisms based on phylogenetic analysis. Ghost lineages assume unseen diversity in the fossil record and serve as predictions for what the fossil record could eventually yield; these hypotheses can be tested by unearthing new fossils or running phylogenetic analyses. Ghost lineages and Lazarus taxa are related concepts, as both stem from gaps in the fossil record. A ghost lineage is any gap in a taxon's fossil record, with or without reappearance, while a Lazarus taxon is a type of ghost lineage wherein a species is believed to have gone extinct due to an absence in the fossil record, but then reappears after a period of time. Exa ...
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