Tapinocephalians
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Tapinocephalians
The Tapinocephalia are one of the major groups of dinocephalian therapsids and the major herbivorous group. Tapinocephalia has been found to consist of three clades: Styracocephalidae, Titanosuchidae, and the very successful Tapinocephalidae. Notable tapinocephalians include ''Moschops'', ''Tapinocephalus'', and ''Titanosuchus''. Description Unlike anteosaurs and estemmenosuchids, tapinocephalians are primarily an African group. The estemmenosuchids and pareiasaurs may have occupied this paleo-bovine niche in the north. Only one tapinocephalian, ''Ulemosaurus'', is known from Russia. Earlier tapinocephalians were carnivorous or omnivorous. One such group was Titanosuchidae, which consisted of long-tailed predators that hunted herbivorous therapsids Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that wer ...
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Titanosuchus
''Titanosuchus ferox'' ("Fierce titan crocodile") is an extinct species of dinocephalian therapsids that lived in the Middle Permian epoch in South Africa. Along with its close relatives, ''Jonkeria'' and ''Moschops'', ''Titanosuchus'' inhabited present-day South Africa around 265 million years ago, in the Late Permian. ''Titanosuchus'' was a carnivore which measured over 2.5 m long and might have eaten both ''Jonkeria'' and ''Moschops'', among other vertebrates. Its teeth included sharp incisors and fang-like canines, perfect for biting prey. ''Titanosuchus'' should not be confused with the therapsid ''Eotitanosuchus'', which belonged to a different family. ''Parascapanodon'' and ''Scapanodon'' were once thought to be distinct genera, but are now considered to be junior synonyms of ''Titanosuchus''.Boonstra, L. D., 1969, The fauna of the Tapinocephalus zone (Beaufort beds of the Karoo): Annals of the South African Museum, v. 56, part 1, p. 1-73. See also * List of therapsids ...
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Tapinocephalians
The Tapinocephalia are one of the major groups of dinocephalian therapsids and the major herbivorous group. Tapinocephalia has been found to consist of three clades: Styracocephalidae, Titanosuchidae, and the very successful Tapinocephalidae. Notable tapinocephalians include ''Moschops'', ''Tapinocephalus'', and ''Titanosuchus''. Description Unlike anteosaurs and estemmenosuchids, tapinocephalians are primarily an African group. The estemmenosuchids and pareiasaurs may have occupied this paleo-bovine niche in the north. Only one tapinocephalian, ''Ulemosaurus'', is known from Russia. Earlier tapinocephalians were carnivorous or omnivorous. One such group was Titanosuchidae, which consisted of long-tailed predators that hunted herbivorous therapsids Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that wer ...
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Moschops
''Moschops'' (Greek for "calf face") is an extinct genus of therapsids that lived in the Guadalupian epoch, around 265–260 million years ago. They were heavily built plant eaters, and they may have lived partly in water, as hippopotamuses do. They had short, thick heads and might have competed by head-butting each other. Their elbow joints allowed them to walk with a more mammal-like gait rather than crawling. Their remains were found in the Karoo region of South Africa, belonging to the ''Tapinocephalus'' Assemblage Zone. Therapsids, such as ''Moschops'', are synapsids, the dominant land animals in the Permian period, which ended 252 million years ago. Description ''Moschops'' were heavy set dinocephalian synapsids, measuring in length, and weighing on average and in maximum body mass. They had small heads with broad orbits and heavily-built short necks. Like other members of Tapinocephalidae, the skull had a tiny opening for the pineal organ. The occiput was broad and ...
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Estemmenosuchidae
Estemmenosuchidae is an extinct family of large, very early herbivorous therapsids that flourished during the Guadalupian period. They are distinguished by horn-like structures, probably for display or agonistic behavior. Apart from the best known genus, '' Estemmenosuchus'', the group is poorly known. To date, their fossils are known only from the Perm region of Russia (a region referred to by Russian paleontologists as the Cis-Urals). Description Estemmenosuchids are among the most distinctive of the Permian tetrapods. The high and massive skull is equipped with a number of horns projecting both upwards and outwards, which were probably used for intra-specific display. The incisors and canine teeth are large, but those at the side are reduced, with a serrated apex, and may have helped to break up plant material, although they were too small to be of much use. The body is large and bulky, indicating a large digestive tract for digesting volumes of plant food. The skull superfi ...
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Styracocephalus
''Styracocephalus platyrhynchus'' ('spike head') is an extinct species of tapinocephalian therapsids that lived during the Guadalupian epoch. ''Styracocephalus''s head ornament meant that it could be recognised from a distance. The most striking feature of ''Styracocephalus'' are the large backward-protruding tabular horns. The crest stuck upwards and backwards, but there is some variation in its shape, and this suggests that it changed throughout life and that it may be sexually dimorphic. ''Styracocephalus'' was a herbivore that may have been fully terrestrial or partly aquatic like the modern hippopotamus. It may have evolved from the estemmenosuchids. Its remains are known from South Africa but it probably had a wider distribution. It was around in length, with a , skull. See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely ve ...
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Tapinocephalidae
Tapinocephalidae was an advanced family of tapinocephalians. It is defined as the clade containing ''Ulemosaurus'', '' Tapinocaninus'', and the Tapinocephalinae. They are known from both Russia and South Africa. In all probability, the Tapinocephalidae had a worldwide (Pangean) distribution. They flourished briefly during the Wordian and Capitanian ages, radiating into several lineages, existing simultaneously, and differing mainly in details of the skull and, to an even lesser degree, the skeleton. It is not clear how such similar animals could each find their own ecological niche, but such was obviously the case. There is a parallel here with the hadrosaur and ceratopsian dinosaurs of the Late Cretaceous. The cause of their abrupt extinction is not clear, since other smaller animals, and even the pareiasaurs, were not affected. Quite probably, like the extinction of the late Pleistocene megafauna, a number of factors were involved. Description The body is deep and capacious, al ...
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Titanosuchidae
Titanosuchidae is an extinct family of dinocephalians. The titanosuchids were carnivorous to omnivorous (herbivorous?) tapinocephalians. As with other tapinocephalians, they had thick skulls probably for head-butting. They had large canine teeth, and their incisors were very strong. They are related to other dinocephalians, such as the Tapinocephalidae - a group that includes ''Moschops''. The most famous titanosuchids are ''Jonkeria'' and ''Titanosuchus ''Titanosuchus ferox'' ("Fierce titan crocodile") is an extinct species of dinocephalian therapsids that lived in the Middle Permian epoch in South Africa. Along with its close relatives, ''Jonkeria'' and ''Moschops'', ''Titanosuchus'' inhabited ...''. References *L. D. Boonstra, "The Fauna of the Tapinocephalus Zone (Beaufort Beds of the Karoo)", Annals of the South African Museum, 56 (1) 1969, pp. 1–73 *Carroll, R. L. Vertebrate paleontology and evolution. -W. H. Freeman and company, New York, 1988 *Edwin H. Colber ...
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Ulemosaurus
''Ulemosaurus'' is an extinct genus of dinocephalian therapsids that lived 265 to 260 million years ago, at Isheevo in Russian Tatarstan. It was a tapinocephalid, a group of bulky herbivores which flourished in the Middle Permian. ''Ulemosaurus'' and other tapinocephalians disappeared at the end of the Middle Permian. Description Only several partial skeletons and skulls have been found. The skull bones are extremely dense: about at its thickest. This thickening is possibly related to head-butting behavior, as some researchers suggest. The species is considered a herbivore, but because the mandible is heavily constructed some palaeontologists consider it a carnivore, with the species being able to use muscle power to cut prey up with its incisors. Classification ''Ulemosaurus'' is a large ''Moschops''-like form from Russia; it is probably similar enough to be included as a separate species of ''Moschops''. Despite its advanced characteristics, it lived slightly before the Karoo ...
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Estemmenosuchid
Estemmenosuchidae is an extinct family of large, very early herbivorous therapsids that flourished during the Guadalupian period. They are distinguished by horn-like structures, probably for display or agonistic behavior. Apart from the best known genus, ''Estemmenosuchus'', the group is poorly known. To date, their fossils are known only from the Perm region of Russia (a region referred to by Russian paleontologists as the Cis-Urals). Description Estemmenosuchids are among the most distinctive of the Permian tetrapods. The high and massive skull is equipped with a number of horns projecting both upwards and outwards, which were probably used for intra-specific display. The incisors and canine teeth are large, but those at the side are reduced, with a serrated apex, and may have helped to break up plant material, although they were too small to be of much use. The body is large and bulky, indicating a large digestive tract for digesting volumes of plant food. The skull superficia ...
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Tapinocephalus
''Tapinocephalus'' ("humble head") is an extinct genus of large herbivorous dinocephalians that lived during the Middle Permian Period. These stocky, barrel-bodied animals were characterised by a massive bony skull roof and short weak snout. It is thought that, like the rest of the members of its family, the animals engaged in head-butting intraspecific behavior, possibly for territory or mates. The fossil remains (skull and postcranial elements) of ''Tapinocephalus'' are known from the Lower, Middle, and Upper part of the ''Tapinocephalus'' Assemblage Zone ( Capitanian age) of the Lower Beaufort Beds of the South African Karoo. Only the type species, ''T. atherstonei'' is now considered valid for this genus. In life, these animals were over in length and massed around , making them among the largest animals of their time. Classification ''Tapinocephalus atherstonei'' is known from a number of skulls and postcranial bones. The skull is large with a heavily pachyostotic ...
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Middle Permian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Omnivorous
An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutrients and energy of the sources absorbed. Often, they have the ability to incorporate food sources such as algae, fungi, and bacteria into their diet. Omnivores come from diverse backgrounds that often independently evolved sophisticated consumption capabilities. For instance, dogs evolved from primarily carnivorous organisms (Carnivora) while pigs evolved from primarily herbivorous organisms (Artiodactyla). Despite this, physical characteristics such as tooth morphology may be reliable indicators of diet in mammals, with such morphological adaptation having been observed in bears. The variety of different animals that are classified as omnivores can be placed into further sub-categories depending on their feeding behaviors. Frugivores ...
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