Tanystropheidae
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Tanystropheidae
Tanystropheidae is an extinct family of mostly marine archosauromorph reptiles that lived throughout the Triassic Period. They are characterized by their long, stiff necks formed from elongated cervical vertebrae with very long cervical ribs. Some tanystropheids such as ''Tanystropheus'' had necks that were several meters long, longer than the rest of their bodies. Tanystropheids are known from Europe, Asia (Russia, China, and Saudi Arabia), North America and probably South America (Brazil). The presence of tanystropheids in Europe and China indicate that they lived along much of the coastline of the Tethys Ocean. However, species in western North America are found in terrestrial deposits, suggesting that as a group, tanystropheids were ecologically diverse. Relationships among tanystropheid species have been difficult to resolve because most specimens were flattened during fossilization and are preserved two-dimensionally. Three-dimensional fossils are known from Europe and N ...
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Ozimek Volans
''Ozimek'' (named after the Ozimek, town of the same name) is a genus of sharovipterygidae, sharovipterygid archosauromorpha, archosauromorph reptile, known from Late Triassic deposits in Poland and closely related to the Kyrgyzstani ''Sharovipteryx''. It contains one species, ''O. volans'', named in 2016 by Jerzy Dzik and Tomasz Sulej. Like ''Sharovipteryx'', ''Ozimek'' had long, slender limbs with the hindlimbs longer than the forelimbs; the hindlimbs likely supported gliding membranes as fossilized in ''Sharovipteryx''. Another unusual characteristic was the shoulder girdle, where the massive coracoids formed a shield-like structure covering the bottom of the shoulder region that would have limited mobility. In other respects, such as its long neck, it was a typical member of the non-natural grouping Protorosauria. Phylogenetic analysis has indicated that it, possibly along with ''Sharovipteryx'', may have been an unusual member of the protorosaur group Tanystropheidae, althoug ...
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Pectodens
''Pectodens'' (meaning "comb tooth") is an extinct genus of archosauromorph reptile which lived during the Middle Triassic in China. The type and only species of the genus is ''P. zhenyuensis'', named by Chun Li and colleagues in 2017. It was a member of the Archosauromorpha, specifically part of the unnatural grouping Protorosauria. However, an unusual combination of traits similar (such as the long neck) and dissimilar (such as the absence of a hook on the fifth metatarsal bone) to other protorosaurs initially led to confusion over its evolutionary relationships. In 2021, it was placed in a newly-established group, Dinocephalosauridae, along with its closest relative '' Dinocephalosaurus''. A small, slender animal measuring long, ''Pectodens'' was named after the peculiar comb-like arrangement of long, conical teeth present in its mouth. Unlike ''Dinocephalosaurus'' and the other reptiles that it was preserved with, well-developed joints and claw-like digits indicate that ''Pec ...
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Protorosaur
Protorosauria is an extinct polyphyletic group of archosauromorph reptiles from the latest Middle Permian (Capitanian stage) to the end of the Late Triassic ( Rhaetian stage) of Asia, Europe and North America. It was named by the English anatomist and paleontologist Thomas Henry Huxley in 1871 as an order, originally to solely contain ''Protorosaurus''. Other names which were once considered equivalent to Protorosauria include Prolacertiformes and Prolacertilia. Protorosaurs are distinguished by their long necks formed by elongated cervical vertebrae, which have ribs that extend backward to the vertebrae behind them. Protorosaurs also have a gap between the quadrate bones and the jugal bones in the back of the skull near the jaw joint, making their skulls resemble those of lizards. While previously thought to be monophyletic, the group is now though to consist of various groups of basal archosauromorph reptiles that lie outside Crocopoda. Classification Protorosauria was ...
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Tanystropheus Longobardicus
''Tanystropheus'' (Greek ~ 'long' + 'hinged') is an extinct archosauromorph reptile from the Middle and Late Triassic epochs. It is recognisable by its extremely elongated neck, which measured long—longer than its body and tail combined. The neck was composed of 12–13 extremely elongated vertebrae. With its very long but relatively stiff neck, ''Tanystropheus'' has been often proposed and reconstructed as an aquatic or semi-aquatic reptile, a theory supported by the fact that the creature is most commonly found in semi-aquatic fossil sites wherein known terrestrial reptile remains are scarce. Fossils have been found in Europe. Complete skeletons of small individuals are common in the Besano Formation at Monte San Giorgio in Italy and Switzerland; other fossils have been found in the Middle East and China, dating from the Middle Triassic to the early part of the Late Triassic (Anisian, Ladinian, and Carnian stages).Dal Sasso, C. and Brillante, G. (2005). ''Dinosaurs of Italy ...
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Archosauromorph
Archosauromorpha (Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, lizards, and snakes). Archosauromorphs first appeared during the late Middle Permian or Late Permian, though they became much more common and diverse during the Triassic period. Although Archosauromorpha was first named in 1946, its membership did not become well-established until the 1980s. Currently Archosauromorpha encompasses four main groups of reptiles: the stocky, herbivorous allokotosaurs and rhynchosaurs, the hugely diverse Archosauriformes, and a polyphyletic grouping of various long-necked reptiles including ''Protorosaurus'', tanystropheids, and ''Prolacerta''. Other groups including pantestudines (turtles and their extinct relatives) and the semiaquatic choristoderes have also been placed in Archosauromorpha by some authors. A ...
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Tanystropheus
''Tanystropheus'' (Greek ~ 'long' + 'hinged') is an extinct archosauromorph reptile from the Middle Triassic, Middle and Late Triassic epochs. It is recognisable by its extremely elongated neck, which measured long—longer than its body and tail combined. The neck was composed of 12–13 extremely elongated vertebrae. With its very long but relatively stiff neck, ''Tanystropheus'' has been often proposed and reconstructed as an aquatic or semi-aquatic reptile, a theory supported by the fact that the creature is most commonly found in semi-aquatic fossil sites wherein known terrestrial reptile remains are scarce. Fossils have been found in Europe. Complete skeletons of small individuals are common in the Besano Formation at Monte San Giorgio in Italy and Switzerland; other fossils have been found in the Middle East and China, dating from the Middle Triassic to the early part of the Late Triassic (Anisian, Ladinian, and Carnian stages).Dal Sasso, C. and Brillante, G. (2005). ''Di ...
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Archosauromorpha
Archosauromorpha (Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, lizards, and snakes). Archosauromorphs first appeared during the late Middle Permian or Late Permian, though they became much more common and diverse during the Triassic period. Although Archosauromorpha was first named in 1946, its membership did not become well-established until the 1980s. Currently Archosauromorpha encompasses four main groups of reptiles: the stocky, herbivorous allokotosaurs and rhynchosaurs, the hugely diverse Archosauriformes, and a polyphyletic grouping of various long-necked reptiles including ''Protorosaurus'', tanystropheids, and ''Prolacerta''. Other groups including pantestudines (turtles and their extinct relatives) and the semiaquatic choristoderes have also been placed in Archosauromorpha by some authors. A ...
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Jesairosaurus
''Jesairosaurus'' is an extinct genus of early archosauromorph reptile known from the Illizi Province of Algeria. It is known from a single species, ''Jesairosaurus lehmani''. Although a potential relative of the long-necked tanystropheids, this lightly-built reptile could instead be characterized by its relatively short neck as well as various skull features. Etymology and discovery Zarzaïtine fossil material has been known since 1957. Much of the material has been recovered by French expeditions in the late 1950s and 1960s, and deposited at the Laboratoire de Paleontologie (Paleontology department) at the Museum national d'Histoire naturalle in Paris. Algerian fossils were prepared at this institution over subsequent years. Several putative procolophonid skeletons reported in 1971 were later determined to belong to " prolacertiforms" in 1990. The term "prolacertiform" is now considered to refer to an unnatural polyphyletic grouping of early archosauromorphs, distant relatives ...
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Dinocephalosauridae
Dinocephalosauridae is an extinct clade of marine and terrestrial archosauromorph reptiles that lived throughout the Triassic period. Like tanystropheids, they are characterized by their long necks, lengthened by either addition of cervical vertebrae or elongation of the individual bones. Dinocephalosaurids are known from Europe (Poland, Germany, Austria, Netherlands) and China. Some members (i.e. '' Dinocephalosaurus'') are solely marine animals, most likely living near the coastlines of the Tethys Ocean, while other members (i.e. ''Pectodens'' were purely terrestrial, suggesting wide ecological diversity in just the few known species in this family. Classification In 2021, a phylogenetic study was conducted by S. Spiekman, N. Fraser, and T. Schayer in an attempt to clarify the systematics of " protorosaur" groups. A total of 16 individual trees were found using different character scoring methods and unstable OTU Otu or OTU may refer to: * Otu: ** Otu, Iran, a village in M ...
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Fuyuansaurus
''Fuyuansaurus'' is an extinct genus of "protorosaur" reptiles (probably part of Tanystropheidae) known from the Middle Triassic (Ladinian stage) Falang Formation of southern China. ''Fuyuansaurus'' was first named by Nicholas C. Fraser, Olivier Rieppel and Li Chun in 2013 and the type species is ''Fuyuansaurus acutirostris''. This genus is known from a single partial skeleton, IVPP V17983. This skeleton includes part of the skull, most of the neck, and portions of the trunk, shoulder girdle, and hip. ''Fuyuansaurus'' can be distinguished from other "protorosaurs" by its elongated and slender snout. Like ''Tanystropheus'', it possessed an elongated neck, but it differs from most tanystropheids in the structure of its hip. Nevertheless, Ezcurra and Butler classified it as a tanystropheid in 2018, as did Spiekman et al. in 2021. Description The skull was crushed and incomplete, with the tip of the snout broken off and the middle part of the skull obscured by the part of the ov ...
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Czatkowiella
''Czatkowiella'' is an extinct genus of long-necked archosauromorph known from Early Triassic (Olenekian age) rocks of Czatkowice 1, Poland. It was first named by Magdalena Borsuk−Białynicka and Susan E. Evans in 2009 and the type species is ''Czatkowiella harae''. Phylogeny Cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... after Borsuk−Białynicka & Evans (2009). Cladogram after Spiekman ''et al.'' 2021: References Prehistoric archosauromorphs Prehistoric reptile genera Olenekian genera Early Triassic reptiles of Europe Fossils of Poland Fossil taxa described in 2009 {{triassic-reptile-stub ...
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Prolacerta
''Prolacerta'' is a genus of archosauromorph from the lower Triassic of South Africa and Antarctica. The only known species is ''Prolacerta broomi''. The generic name ''Prolacerta'' is derived from Latin meaning “before lizard” and its species name ''broomi'' is in commemoration of the famous paleontologist Robert Broom, who discovered and studied many of the fossils found in rocks of the Karoo Supergroup. When first discovered, ''Prolacerta'' was considered to be ancestral to modern lizards, scientifically known as lacertilians. However, a study by Gow (1975) instead found that it shared more similarities with the lineage that would lead to archosaurs such as crocodilians and dinosaurs (including birds). ''Prolacerta'' is considered by modern paleontologists to be among the closest relatives of the Archosauriformes. History of discovery ''Prolacerta'' was first described by Francis Rex Parrington in 1935 from a single skull recovered near the small town, Middelburg, in the ...
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