Tylomelania Toradjarum
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Tylomelania Toradjarum
''Tylomelania toradjarum'' is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Pachychilidae. The specific name ''toradjarum'' is in honor of Toraja (spelling variant Toradja), an ethnic group indigenous to Sulawesi. Distribution This species is endemic to Lake Poso, Sulawesi, Indonesia. The type locality is the Lake Poso. Description The shell has 7-9 whorls A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). Whorls in nature File:Photograph and axial plane floral d .... The width of the shell is 16 mm. The height of the shell is 54 mm. The width of the aperture is 10 mm. The height of the aperture is 15 mm. There are 9 concentric lines on the operculum. References External links * von Rintelen T. & Glaubrecht M. (2005). "Anatomy of an adaptive radiation: a ...
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Gastropod Shell
The gastropod shell is part of the body of a Gastropoda, gastropod or snail, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some gastropods appear shell-less (slugs) but may have a remnant within the mantle, or in some cases the shell is reduced such that the body cannot be retracted within it (semi-slug). Some snails also possess an operculum that seals the opening of the shell, known as the Aperture (mollusc), aperture, which provides further protection. The study of mollusc shells is known as conchology. The biological study of gastropods, and other molluscs in general, is malacology. Shell morphology terms vary by species group. Shell layers The gastropod shell has three major layers secreted by the Mantle (mollusc), mantle. The calcareous central layer, tracum, is typically made of calcium carbonate precipitated into an organic matrix known as c ...
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Specific Name (zoology)
In zoological nomenclature, the specific name (also specific epithet or species epithet) is the second part (the second name) within the scientific name of a species (a binomen). The first part of the name of a species is the name of the genus or the generic name. The rules and regulations governing the giving of a new species name are explained in the article species description. For example, the scientific name for humans is ''Homo sapiens'', which is the species name, consisting of two names: ''Homo'' is the " generic name" (the name of the genus) and ''sapiens'' is the "specific name". Historically, ''specific name'' referred to the combination of what are now called the generic and specific names. Carl Linnaeus, who formalized binomial nomenclature, made explicit distinctions between specific, generic, and trivial names. The generic name was that of the genus, the first in the binomial, the trivial name was the second name in the binomial, and the specific the proper term for ...
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Biological Journal Of The Linnean Society
The ''Biological Journal of the Linnean Society'' is a direct descendant of the oldest biological journal in the world, the ''Transactions of the Linnean Society''. It succeeded the earlier title in 1969. The journal specializes in evolution, and encompasses work across all taxonomic groups in all five kingdoms of living organisms. It includes all methods, whole-organism or molecular, practical or theoretical. The journal is published by the Linnean Society of London The Linnean Society of London is a learned society dedicated to the study and dissemination of information concerning natural history, evolution, and taxonomy. It possesses several important biological specimen, manuscript and literature colle .... References External links * Linnean Society of London Biology journals English-language journals Publications with year of establishment missing {{biology-journal-stub ...
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Operculum (gastropod)
The operculum (; ) is a corneous or calcareous anatomical structure like a trapdoor that exists in many (but not all) groups of sea snails and freshwater snails, and also in a few groups of land snails; the structure is found in some marine and freshwater gastropods, and in a minority of terrestrial gastropods, including the families Helicinidae, Cyclophoridae, Aciculidae, Maizaniidae, Pomatiidae, etc. The operculum is attached to the upper surface of the foot and in its most complete state, it serves as a sort of "trapdoor" to close the aperture of the shell when the soft parts of the animal are retracted. The shape of the operculum varies greatly from one family of gastropods to another. It is fairly often circular, or more or less oval in shape. In species where the operculum fits snugly, its outline corresponds exactly to the shape of the aperture of the shell and it serves to seal the entrance of the shell. Many families have opercula that are reduced in size, and which a ...
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Whorl (mollusc)
A whorl is a single, complete 360° revolution or turn in the spiral growth of a mollusc shell. A spiral configuration of the shell is found in numerous gastropods, but it is also found in shelled cephalopods including ''Nautilus'', ''Spirula'' and the large extinct subclass of cephalopods known as the ammonites. A spiral shell can be visualized as consisting of a long conical tube, the growth of which is coiled into an overall helical or planispiral shape, for reasons of both strength and compactness. The number of whorls which exist in an adult shell of a particular species depends on mathematical factors in the geometric growth, as described in D'Arcy Wentworth Thompson's classic 1917 book ''On Growth and Form'', and by David Raup. The main factor is how rapidly the conical tube expands (or flares-out) over time. When the rate of expansion is low, such that each subsequent whorl is not that much wider than the previous one, then the adult shell has numerous whorls. When the ...
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Naturalis Biodiversity Center - RMNH
Naturalis Biodiversity Center ( nl, Nederlands Centrum voor Biodiversiteit Naturalis) is a national museum of natural history and a research center on biodiversity in Leiden, Netherlands. It was named the European Museum of the Year 2021. Although its current name and organization are relatively recent, the history of Naturalis can be traced back to the early 1800s. Its collection includes approximately 42 million specimens, making it one of the largest natural history collections in the world. History The beginnings of Naturalis go back to the creation of the Rijksmuseum van Natuurlijke Historie (abbreviated RMNH, National Museum of Natural History) by Dutch King William I on August 9, 1820. In 1878, the geological and mineralogical collections of the museum were split off into a separate museum, remaining distinct until the merger of the Rijksmuseum van Natuurlijke Historie with the Rijksmuseum van Geologie en Mineralogie (abbreviated RGM) in 1984, to form the Nationaal Natu ...
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Type Locality (biology)
In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular taxon. In older usage (pre-1900 in botany), a type was a taxon rather than a specimen. A taxon is a scientifically named grouping of organisms with other like organisms, a set that includes some organisms and excludes others, based on a detailed published description (for example a species description) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution. Type specimen According to a precise set of rules laid down in the International Code of Zoological Nomenclature (ICZN) and the International Code of Nomenclature for algae, fungi, and plants (ICN), the scientific name of every taxon is almost a ...
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Lake Poso
Lake Poso ( id, Danau Poso) is a lake in Central Sulawesi, Indonesia, and the third-deepest lake in Indonesia. The town of Pendolo is situated at the southern end of the lake, the town of Tentena is located at the northern end, while a number of smaller villages dot the shoreline. The lake drains into the Poso River at Tentena, which flows into the Molucca Sea at the town of Poso. Ecology The lake contains various fish, including the eel '' Anguilla marmorata'' which migrates between the lake and the sea, and 11 fish species that are endemic to the lake, notably buntingi ricefish ('' Adrianichthys'', '' Oryzias nebulosus'', '' O. nigrimas'' and '' O. orthognathus''), gobies ('' Mugilogobius amadi'' and '' M. sarasinorum''), and the halfbeak '' Nomorhamphus celebensis''. These endemics are all highly threatened; in some cases possibly already extinct. One of the reasons for the drastic decline of the native fish are introduced, non-native species, particularly Mozambique ti ...
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Sulawesi
Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Archipelago. Within Indonesia, only Sumatra, Borneo, and New Guinea, Papua are larger in territory, and only Java and Sumatra have larger populations. The landmass of Sulawesi includes four peninsulas: the northern Minahassa Peninsula, Minahasa Peninsula, the East Peninsula, Sulawesi, East Peninsula, the South Peninsula, Sulawesi, South Peninsula, and the Southeast Peninsula, Sulawesi, Southeast Peninsula. Three gulfs separate these peninsulas: the Gulf of Tomini between the northern Minahasa and East peninsulas, the Tolo Gulf between the East and Southeast peninsulas, and the Bone Gulf between the South and Southeast peninsulas. The Strait of Makassar runs along the western side of the island and separates the island from Borneo. Etymology ...
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Toraja
The Torajans are an ethnic group indigenous to a mountainous region of South Sulawesi, Indonesia. Their population is approximately 1,100,000, of whom 450,000 live in the regency of Tana Toraja ("Land of Toraja"). Most of the population is Christian, and others are Muslim or have local animist beliefs known as ''aluk'' ("the way"). The Indonesian government has recognised this animistic belief as ''Aluk To Dolo'' ("Way of the Ancestors"). The word ''Toraja'' comes from the Buginese language term ''to riaja'', meaning "people of the uplands". The Dutch colonial government named the people ''Toraja'' in 1909. Torajans are renowned for their elaborate funeral rites, burial sites carved into rocky cliffs, massive peaked-roof traditional houses known as ''tongkonan'', and colourful wood carvings. Toraja funeral rites are important social events, usually attended by hundreds of people and lasting for several days. Before the 20th century, Torajans lived in autonomous villages, where ...
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Pachychilidae
Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.Gofas, S. (2014). Pachychilidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=411651 on 2014-11-20 Distribution Pachychilids are freshwater snails with a worldwide distribution in the tropics. Representatives are found in South and Central America, Africa, Madagascar, South and South-east Asia and tropical Australia (Queensland: Torres Strait Islands). Description Pachychilids have an operculum, which is concentric and multispiral. Ecology All species in the family inhabit freshwater except '' Faunus ater'', which is a brackish water snail found in estuaries and other coastal habitats. Pachychilids are either oviparous (lay eggs), ovoviviparous or viviparous (retain developing eggs and youngs in special incubatory structures). Notes on the taxonomy The name is derived from a co ...
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Paul Sarasin
Paul Sarasin, full name Paul Benedict Sarasin (11 December 1856 – 7 April 1929) was a Swiss people, Swiss naturalist and ethnologist. He is known as founder of National parks in Switzerland.Bibliography of Paul Sarasin
ETH-Bibliothek Zürich, accessed 9 August 2010.


Life and work

Paul Sarasin studied medicine and natural science at the University of Basel with Leopold Ruetimeyer, where he also met Fritz Sarasin, and at the University of Würzburg. His dissertation had the title "Die Entwicklungsgeschichte der Bithynia (gastropod), Bithynia tentaculata" and was about the developmental history of a small aquatic snail. Paul Sarasin was a second cousin of Fritz Sarasin.Coan E. V., Kabat A. R. & Petit R. E. (15 February 2 ...
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