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Tylomelania Carbo
''Tylomelania carbo'' is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Pachychilidae. The sister group (the closest relative) of ''Tylomelania carbo'' is ''Tylomelania toradjarum''. Distribution This species occurs in lake Poso, Central Sulawesi, Indonesia. The type locality is lake Poso. Description The shell has 4-5 whorls. The width of the shell is up to 10.5 mm. The height of the shell is up to 16.5 mm. The width of the aperture is 6 mm. The height of the aperture is 9 mm. The length of the radula can be up to 21.5 mm.von Rintelen T. & Glaubrecht M. (2005). "Anatomy of an adaptive radiation: a unique reproductive strategy in the endemic freshwater gastropod ''Tylomelania'' (Cerithioidea: Pachychilidae) on Sulawesi, Indonesia and its biogeographical implications." '' Biological Journal of the Linnean Society'' 85: 513–542. . References External links * Sarasin P. & Sarasin F. (1898). ...
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Gastropod Shell
The gastropod shell is part of the body of a Gastropoda, gastropod or snail, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some gastropods appear shell-less (slugs) but may have a remnant within the mantle, or in some cases the shell is reduced such that the body cannot be retracted within it (semi-slug). Some snails also possess an operculum that seals the opening of the shell, known as the Aperture (mollusc), aperture, which provides further protection. The study of mollusc shells is known as conchology. The biological study of gastropods, and other molluscs in general, is malacology. Shell morphology terms vary by species group. Shell layers The gastropod shell has three major layers secreted by the Mantle (mollusc), mantle. The calcareous central layer, tracum, is typically made of calcium carbonate precipitated into an organic matrix known as c ...
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Pachychilidae
Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.Gofas, S. (2014). Pachychilidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=411651 on 2014-11-20 Distribution Pachychilids are freshwater snails with a worldwide distribution in the tropics. Representatives are found in South and Central America, Africa, Madagascar, South and South-east Asia and tropical Australia (Queensland: Torres Strait Islands). Description Pachychilids have an operculum, which is concentric and multispiral. Ecology All species in the family inhabit freshwater except '' Faunus ater'', which is a brackish water snail found in estuaries and other coastal habitats. Pachychilids are either oviparous (lay eggs), ovoviviparous or viviparous (retain developing eggs and youngs in special incubatory structures). Notes on the taxonomy The name is derived from a co ...
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Biological Journal Of The Linnean Society
The ''Biological Journal of the Linnean Society'' is a direct descendant of the oldest biological journal in the world, the ''Transactions of the Linnean Society''. It succeeded the earlier title in 1969. The journal specializes in evolution, and encompasses work across all taxonomic groups in all five kingdoms of living organisms. It includes all methods, whole-organism or molecular, practical or theoretical. The journal is published by the Linnean Society of London The Linnean Society of London is a learned society dedicated to the study and dissemination of information concerning natural history, evolution, and taxonomy. It possesses several important biological specimen, manuscript and literature colle .... References External links * Linnean Society of London Biology journals English-language journals Publications with year of establishment missing {{biology-journal-stub ...
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Radula
The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters the esophagus. The radula is unique to the molluscs, and is found in every class of mollusc except the bivalves, which instead use cilia, waving filaments that bring minute organisms to the mouth. Within the gastropods, the radula is used in feeding by both herbivorous and carnivorous snails and slugs. The arrangement of teeth ( denticles) on the radular ribbon varies considerably from one group to another. In most of the more ancient lineages of gastropods, the radula is used to graze, by scraping diatoms and other microscopic algae off rock surfaces and other substrates. Predatory marine snails such as the Naticidae use the radula plus an acidic secretion to bore through the shell of other molluscs. Other predatory marine snails ...
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Whorl (mollusc)
A whorl is a single, complete 360° revolution or turn in the spiral growth of a mollusc shell. A spiral configuration of the shell is found in numerous gastropods, but it is also found in shelled cephalopods including ''Nautilus'', ''Spirula'' and the large extinct subclass of cephalopods known as the ammonites. A spiral shell can be visualized as consisting of a long conical tube, the growth of which is coiled into an overall helical or planispiral shape, for reasons of both strength and compactness. The number of whorls which exist in an adult shell of a particular species depends on mathematical factors in the geometric growth, as described in D'Arcy Wentworth Thompson's classic 1917 book ''On Growth and Form'', and by David Raup. The main factor is how rapidly the conical tube expands (or flares-out) over time. When the rate of expansion is low, such that each subsequent whorl is not that much wider than the previous one, then the adult shell has numerous whorls. When the ...
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Type Locality (biology)
In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular taxon. In older usage (pre-1900 in botany), a type was a taxon rather than a specimen. A taxon is a scientifically named grouping of organisms with other like organisms, a set that includes some organisms and excludes others, based on a detailed published description (for example a species description) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution. Type specimen According to a precise set of rules laid down in the International Code of Zoological Nomenclature (ICZN) and the International Code of Nomenclature for algae, fungi, and plants (ICN), the scientific name of every taxon is almost a ...
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Lake Poso
Lake Poso ( id, Danau Poso) is a lake in Central Sulawesi, Indonesia, and the third-deepest lake in Indonesia. The town of Pendolo is situated at the southern end of the lake, the town of Tentena is located at the northern end, while a number of smaller villages dot the shoreline. The lake drains into the Poso River at Tentena, which flows into the Molucca Sea at the town of Poso. Ecology The lake contains various fish, including the eel ''Anguilla marmorata'' which migrates between the lake and the sea, and 11 fish species that are endemic to the lake, notably buntingi ricefish ('' Adrianichthys'', ''Oryzias nebulosus'', '' O. nigrimas'' and '' O. orthognathus''), gobies (''Mugilogobius amadi'' and '' M. sarasinorum''), and the halfbeak ''Nomorhamphus celebensis''. These endemics are all highly threatened; in some cases possibly already extinct. One of the reasons for the drastic decline of the native fish are introduced, non-native species, particularly Mozambique t ...
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Tylomelania Toradjarum
''Tylomelania toradjarum'' is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Pachychilidae. The specific name ''toradjarum'' is in honor of Toraja (spelling variant Toradja), an ethnic group indigenous to Sulawesi. Distribution This species is endemic to Lake Poso, Sulawesi, Indonesia. The type locality is the Lake Poso. Description The shell has 7-9 whorls A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). Whorls in nature File:Photograph and axial plane floral d .... The width of the shell is 16 mm. The height of the shell is 54 mm. The width of the aperture is 10 mm. The height of the aperture is 15 mm. There are 9 concentric lines on the operculum. References External links * von Rintelen T. & Glaubrecht M. (2005). "Anatomy of an adaptive radiation: a ...
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Sister Group
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomic ...
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Mollusk
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gas ...
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Paul Sarasin
Paul Sarasin, full name Paul Benedict Sarasin (11 December 1856 – 7 April 1929) was a Swiss people, Swiss naturalist and ethnologist. He is known as founder of National parks in Switzerland.Bibliography of Paul Sarasin
ETH-Bibliothek Zürich, accessed 9 August 2010.


Life and work

Paul Sarasin studied medicine and natural science at the University of Basel with Leopold Ruetimeyer, where he also met Fritz Sarasin, and at the University of Würzburg. His dissertation had the title "Die Entwicklungsgeschichte der Bithynia (gastropod), Bithynia tentaculata" and was about the developmental history of a small aquatic snail. Paul Sarasin was a second cousin of Fritz Sarasin.Coan E. V., Kabat A. R. & Petit R. E. (15 February 2 ...
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Gastropod
The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. There are many thousands of species of sea snails and slugs, as well as freshwater snails, freshwater limpets, and land snails and slugs. The class Gastropoda contains a vast total of named species, second only to the insects in overall number. The fossil history of this class goes back to the Late Cambrian. , 721 families of gastropods are known, of which 245 are extinct and appear only in the fossil record, while 476 are currently extant with or without a fossil record. Gastropoda (previously known as univalves and sometimes spelled "Gasteropoda") are a major part of the phylum Mollusca, and are the most highly diversified class in the phylum, with 65,000 to 80,000 living snail and slug species. The anatomy, behavior, feeding, and re ...
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