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Tralkasaurus Cuyi
''Tralkasaurus'' (meaning "thunder lizard") is a genus of abelisaurid dinosaur from the Huincul Formation from Río Negro Province in Argentina. The type and only species is ''Tralkasaurus cuyi'', named in 2020 by Mauricio Cerroni and colleagues based on an incomplete skeleton. A medium-sized abelisaurid, ''Tralkasaurus'' exhibits a conflicting blend of characteristics found among the early-diverging abelisauroids with others that characterize the highly specialized clade Brachyrostra, and thus its position within the clade is poorly-resolved. Discovery and naming The Violante Farm fossil site in Río Negro Province, Argentina, near the village of El Cuy, was discovered in 1999 by Sebastián Apesteguía on the north shore of the Ezequiel Ramos Mexía Reservoir. In a 2018 conference abstract, Matías Motta and colleagues reported new fossil discoveries, including a diverse fauna of dinosaurs, made at the Violante Farm site. The strata exposed are yellowish and greenish sandstone ...
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Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding age. Both age and stage bear the same name. As a unit of geologic time measure, the Cenomanian Age spans the time between 100.5 and 93.9 million years ago (Mya). In the geologic timescale, it is preceded by the Albian and is followed by the Turonian. The Upper Cenomanian starts around at 95 Mya. The Cenomanian is coeval with the Woodbinian of the regional timescale of the Gulf of Mexico and the early part of the Eaglefordian of the regional timescale of the East Coast of the United States. At the end of the Cenomanian, an anoxic event took place, called the Cenomanian-Turonian boundary event or the "Bonarelli event", that is associated with a minor extinction event for marine spec ...
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Abelisauridae
Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are found on the modern continents of Africa and South America, as well as on the Indian subcontinent and the island of Madagascar. Isolated teeth were found in the Late Jurassic of Portugal, and the Late Cretaceous genera '' Tarascosaurus'' and ''Arcovenator'' have been described in France. Abelisaurids first appear in the fossil record of the early middle Jurassic period, and at least two genera (the Moroccan ''Chenanisaurus'' and the Madagascan ''Majungasaurus'') survived until the end of the Mesozoic era 66 million years ago. Like most theropods, abelisaurids were carnivorous bipeds. They were characterized by stocky hind limbs and extensive ornamentation of the skull bones, with grooves and pits. In many abelisaurids, such as ''Carnotaurus'', ...
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Carnotaurus
''Carnotaurus'' (; ) is a genus of theropod dinosaur that lived in South America during the Late Cretaceous period, probably sometime between 71 and 69 million years ago. The only species is ''Carnotaurus sastrei''. Known from a single well-preserved skeleton, it is one of the best-understood theropods from the Southern Hemisphere. The skeleton, found in 1984, was uncovered in the Chubut Province of Argentina from rocks of the La Colonia Formation. ''Carnotaurus'' is a derived member of the Abelisauridae, a group of large theropods that occupied the large predatorial niche in the southern landmasses of Gondwana during the late Cretaceous. Within the Abelisauridae, the genus is often considered a member of the Brachyrostra, a clade of short-snouted forms restricted to South America. ''Carnotaurus'' was a lightly built, bipedal predator, measuring in length and weighing . As a theropod, ''Carnotaurus'' was highly specialized and distinctive. It had thick horns above the eyes, ...
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Averostra
Averostra, or "bird snouts", is a clade that includes most theropod dinosaurs, namely Ceratosauria and Tetanurae, and represent the only group of post-Early Jurassic theropods. Both survived into the Cretaceous period. When the Cretaceous–Paleogene extinction event occurred, ceratosaurians, megaraptorans an ''incertae sedis'' group within Tetanurae, and two groups of tetanurans within the clade Coelurosauria, the Tyrannosauroidea and Maniraptoriformes, were still extant. Only one subgroup of Maniraptoriformes, Aves, survived the extinction event and persisted to the present day. One important diagnostic feature of Averostra is the absence of the fifth metacarpal. Other saurischians retained this bone, albeit in a significantly reduced form. Definition Averostra was named by Gregory S. Paul in 2002 as an apomorphy-based clade defined as the group including the Dromaeosauridae and other Avepoda with (an ancestor with) a promaxillary fenestra (''fenestra promaxillaris''), an extra ...
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Masiakasaurus
''Masiakasaurus'' is a genus of small predatory noasaurid theropod dinosaurs from the Late Cretaceous of Madagascar. In Malagasy, ''masiaka'' means "vicious"; thus, the genus name means "vicious lizard". The type species, ''Masiakasaurus knopfleri'', was named after the musician Mark Knopfler, whose music inspired the expedition crew. It was named in 2001 by Scott D. Sampson, Matthew Carrano, and Catherine A. Forster. Unlike most theropods, the front teeth of ''M. knopfleri'' projected forward instead of straight down. This unique dentition suggests that they had a specialized diet, perhaps including fish and other small prey. Other bones of the skeleton indicate that ''Masiakasaurus'' were bipedal, with much shorter forelimbs than hindlimbs. ''M. knopfleri'' was a small theropod, reaching long and weighing . ''Masiakasaurus'' lived around 70 million years ago, along with animals such as ''Majungasaurus'', ''Rapetosaurus'', and ''Rahonavis''. ''Masiakasaurus'' was a member o ...
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Noasaurus
''Noasaurus'' ("Northwestern Argentina lizard") is a genus of ceratosaurian theropod dinosaur genus from the late Campanian-Maastrichtian (Late Cretaceous) of Argentina. The type and only species is ''N. leali''. Discovery and naming In the mid-seventies, a fragmentary small theropod skeleton was discovered by Jaime Eduardo Powell and José Fernando Bonaparte at the Estancia El Brete-site. In 1977, the discovery was reported in the scientific literature. The type species, ''Noasaurus leali'', was named and described by Bonaparte and Powell in 1980. The generic name begins with a usual abbreviation of ''noroeste Argentina'', "northwest Argentina". The specific name honours the owner of the site, Fidel Leal.J. F. Bonaparte and J. E. Powell. 1980. "A continental assemblage of tetrapods from the Upper Cretaceous beds of El Brete, northwestern Argentina (Sauropoda-Coelurosauria-Carnosauria-Aves)". ''Mémoires de la Société Géologique de France, Nouvelle Série'' 139: 19-28 Th ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Candeleros Formation
The Candeleros Formation (formerly known as the Candeleros Member of the "Río Limay Formation") is a geologic formation that crops out in the Río Negro, Neuquén, and Mendoza provinces of northern Patagonia, Argentina. It is the oldest formation in the Neuquén Group and belongs to the Rio Limay Subgroup. Formerly that subgroup was treated as a formation, and the Candeleros Formation was known as the Candeleros Member.Sánchez ''et al.'', 2006 Description The type locality of the Candeleros Formation is Candeleros Hill in Neuquén Province, after which the formation was named by Wichmann in 1929.Wichmann, 1929 This formation unconformably overlies the Lohan Cura Formation, and it is in turn overlain by the Huincul Formation, also a unit of the Neuquén Group. The sediments of the latter are of lighter greenish and yellow colors and the boundary between the Candeleros and Huincul formations is easily recognizable.Leanza ''et al.'', 2004 The Candeleros Formation is almost th ...
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Skorpiovenator
''Skorpiovenator'' ("scorpion hunter") is a genus of abelisaurid theropod dinosaur from the Late Cretaceous (Cenomanian to Turonian) Huincul Formation of Argentina. It is one of the most complete and informative abelisaurids yet known, described from a nearly complete and articulated skeleton. Description The preserved length of the excavated ''Skorpiovenator'' skeleton from the premaxilla to the 12th caudal vertebra is . It was estimated to have grown up to in length. In 2010, Gregory S. Paul gave larger estimations of and .Paul, G.S. (2010) ''The Princeton Field Guide to Dinosaurs'', Princeton University Press p. 81 In 2016, a similar size to the original estimate at has been estimated. It had short, stubby, near-useless arms, but strong legs with powerful thighs and sturdy shins over which its large body was balanced. Skull ''Skorpiovenators skull was short, stout and covered in the ridges, furrows, tubercles and bumpy nodules that are scattered over the heads of most ...
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Mapudungun
Mapuche (, Mapuche & Spanish: , or Mapudungun; from ' 'land' and ' 'speak, speech') is an Araucanian language related to Huilliche spoken in south-central Chile and west-central Argentina by the Mapuche people (from ''mapu'' 'land' and ''che'' 'people'). It is also spelled Mapuzugun and Mapudungu. It was formerly known as Araucanian, the name given to the Mapuche by the Spaniards; the Mapuche avoid it as a remnant of Spanish colonialism. Mapudungun is not an official language of the countries Chile and Argentina, receiving virtually no government support throughout its history. However, since 2013, Mapuche, along with Spanish, has been granted the status of an official language by the local government of Galvarino, one of the many Communes of Chile. It is not used as a language of instruction in either country's educational system despite the Chilean government's commitment to provide full access to education in Mapuche areas in southern Chile. There is an ongoing political ...
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Binomial Nomenclature
In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomial name (which may be shortened to just "binomial"), a binomen, name or a scientific name; more informally it is also historically called a Latin name. The first part of the name – the '' generic name'' – identifies the genus to which the species belongs, whereas the second part – the specific name or specific epithet – distinguishes the species within the genus. For example, modern humans belong to the genus ''Homo'' and within this genus to the species ''Homo sapiens''. ''Tyrannosaurus rex'' is likely the most widely known binomial. The ''formal'' introduction of this system of naming species is credit ...
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Theropoda
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic Period (geology), period 231.4 million years ago (Year#SI prefix multipliers, Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of di ...
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