Sphaerotholus Goodwini
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Sphaerotholus Goodwini
''Sphaerotholus'' is a genus of pachycephalosaurid dinosaur from the Upper Cretaceous of the western United States and Canada. To date, three species have been described: the type species, ''S. goodwini'', from the Den-na-zin Member of the Kirtland Formation (Late Campanian) of San Juan County, New Mexico, USA; ''S. buchholtzae'', from the Hell Creek Formation (Late Maastrichtian) of western Carter County, Montana, USA and the Frenchman Formation of Saskatchewan, Canada; and ''S. edmontonensis'', from the Horseshoe Canyon Formation of Alberta, Canada. History of discovery The etymology of ''Sphaerotholus'' is a combination of the Greek ''sphaira'', meaning "ball", and ''tholos'', meaning "dome", and is a reference to the characteristically dome-shaped pachycephalosaurian skull. The survival of ''Sphaerotholus'' from the Campanian of New Mexico to the end of the Maastrichtian of Montana demonstrates that this taxon had both a relatively long duration (approximately 7-8 mil ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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Frenchman Formation
The Frenchman Formation is stratigraphic unit of Late Cretaceous (late Maastrichtian) age in the Western Canada Sedimentary Basin. It is present in southern Saskatchewan and the Cypress Hills of southeastern Alberta. The formation was defined by G.M. Furnival in 1942 from observations of outcrops along the Frenchman River, between Ravenscrag and Highway 37. It contains the youngest of dinosaur genera, much like the Hell Creek Formation in the United States. Lithology The Frenchman Formation consists of olive-green to brown, fine- to coarse-grained, cross-bedded sandstone with interbedded claystone bands and minor beds and lenses of intraformational clay-clast conglomerate. A conglomerate layer with well-rounded quartzite pebbles is present above the basal unconformity in some areas.Glass, D.J., editor, 1997. Lexicon of Canadian Stratigraphy, vol. 4, Western Canada. Canadian Society of Petroleum Geologists, Calgary, Alberta, 1423 p. on CD-ROM, . Thickness and distribution The ...
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Squamosal
The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal The quadratojugal is a skull bone present in many vertebrates, including some living reptiles and amphibians. Anatomy and function In animals with a quadratojugal bone, it is typically found connected to the jugal (cheek) bone from the front and ... bones form the cheek series of the skull. The bone forms an ancestral component of the dermal roof and is typically thin compared to other skull bones. The squamosal bone lies Anatomical terms of location, ventral to the temporal series and otic notch, and is bordered anteriorly by the Postorbital bone, postorbital. Posteriorly, the squamosal articulates with the quadrate bone, quadrate and Pterygoid bone, pterygoid bones. The squamosal is bordered anteroventrally by the jugal and ventrally by the quadratojugal. Function in reptiles In reptiles, the Quadrate ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Anatomical Terms Of Location
Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provides a definition of what is at the front ("anterior"), behind ("posterior") and so on. As part of defining and describing terms, the body is described through the use of anatomical planes and anatomical axes. The meaning of terms that are used can change depending on whether an organism is bipedal or quadrupedal. Additionally, for some animals such as invertebrates, some terms may not have any meaning at all; for example, an animal that is radially symmetrical will have no anterior surface, but can still have a description that a part is close to the middle ("proximal") or further from the middle ("distal"). International organisations have determined vocabularies that are often used as standard vocabularies for subdisciplines of anatom ...
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Taxon
In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and given a particular ranking, especially if and when it is accepted or becomes established. It is very common, however, for taxonomists to remain at odds over what belongs to a taxon and the criteria used for inclusion. If a taxon is given a formal scientific name, its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping. Initial attempts at classifying and ordering organisms (plants and animals) were set forth in Carl Linnaeus's Linnaean taxonomy, system in ''Systema Naturae'', 10th edition (1758), as well as an unpublished work by Bernard de Jussieu, Bernard and Antoine Laurent de Jussieu. The idea of a unit-based system of biological classification was first mad ...
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Skull (anatomy)
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, these two parts are the neurocranium and the viscerocranium ( facial skeleton) that includes the mandible as its largest bone. The skull forms the anterior-most portion of the skeleton and is a product of cephalisation—housing the brain, and several sensory structures such as the eyes, ears, nose, and mouth. In humans these sensory structures are part of the facial skeleton. Functions of the skull include protection of the brain, fixing the distance between the eyes to allow stereoscopic vision, and fixing the position of the ears to enable sound localisation of the direction and distance of sounds. In some animals, such as horned ungulates (mammals with hooves), the skull also has a defensive function by providing the mount (on the fro ...
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Greek Language
Greek ( el, label=Modern Greek, Ελληνικά, Elliniká, ; grc, Ἑλληνική, Hellēnikḗ) is an independent branch of the Indo-European family of languages, native to Greece, Cyprus, southern Italy (Calabria and Salento), southern Albania, and other regions of the Balkans, the Black Sea coast, Asia Minor, and the Eastern Mediterranean. It has the longest documented history of any Indo-European language, spanning at least 3,400 years of written records. Its writing system is the Greek alphabet, which has been used for approximately 2,800 years; previously, Greek was recorded in writing systems such as Linear B and the Cypriot syllabary. The alphabet arose from the Phoenician script and was in turn the basis of the Latin, Cyrillic, Armenian, Coptic, Gothic, and many other writing systems. The Greek language holds a very important place in the history of the Western world. Beginning with the epics of Homer, ancient Greek literature includes many works of lasting impo ...
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Etymology
Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the Phonological change, form of words and, by extension, the origin and evolution of their semantic meaning across time. It is a subfield of historical linguistics, and draws upon comparative semantics, Morphology_(linguistics), morphology, semiotics, and phonetics. For languages with a long recorded history, written history, etymologists make use of texts, and texts about the language, to gather knowledge about how words were used during earlier periods, how they developed in Semantics, meaning and Phonological change, form, or when and how they Loanword, entered the language. Etymologists also apply the methods of comparative linguistics to reconstruct information about forms that are too old for any direct information to be available. By analyzing related ...
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Sphaerotholus
''Sphaerotholus'' is a genus of pachycephalosaurid dinosaur from the Upper Cretaceous of the western United States and Canada. To date, three species have been described: the type species, ''S. goodwini'', from the Den-na-zin Member of the Kirtland Formation (Late Campanian) of San Juan County, New Mexico, San Juan County, New Mexico, USA; ''S. buchholtzae'', from the Hell Creek Formation (Late Maastrichtian) of western Carter County, Montana, Carter County, Montana, USA and the Frenchman Formation of Saskatchewan, Canada; and ''S. edmontonensis'', from the Horseshoe Canyon Formation of Alberta, Canada. History of discovery The etymology of ''Sphaerotholus'' is a combination of the Greek language, Greek ''sphaira'', meaning "ball", and ''tholos'', meaning "dome", and is a reference to the characteristically dome-shaped pachycephalosaurian skull (anatomy), skull. The survival of ''Sphaerotholus'' from the Campanian of New Mexico to the end of the Maastrichtian of Montana demonstr ...
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