Silurian-Devonian Terrestrial Revolution
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Silurian-Devonian Terrestrial Revolution
The Silurian-Devonian Terrestrial Revolution, also known as the Devonian Plant Explosion (DePE) and the Devonian explosion, was a period of rapid plant and fungal diversification that occurred 428 to 359 million years ago during the Silurian and Devonian, with the most critical phase occurring during the Late Silurian and Early Devonian. This diversification of terrestrial plant life had vast impacts on the biotic composition of earth's soil, its atmosphere, its oceans, and for all plant and animal life that would follow it. Through fierce competition for light and available space on land, phenotypic diversity of plants increased greatly, comparable in scale and effect to the explosion in diversity of animal life during the Cambrian explosion, especially in vertical plant growth, which allowed for photoautotrophic canopies to develop, and forever altering plant evolutionary floras that followed. This Silurian and Devonian flora was significantly different in appearance, reproduction ...
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Arbuscular Mycorrhiza
An arbuscular mycorrhiza (AM) (plural ''mycorrhizae'', a.k.a. ''endomycorrhiza'') is a type of mycorrhiza in which the symbiont fungus (''AM fungi'', or AMF) penetrates the cortical cells of the roots of a vascular plant forming arbuscules. (Not to be confused with ectomycorrhiza or ericoid mycorrhiza.) Arbuscular mycorrhizae are characterized by the formation of unique structures, arbuscules and vesicles, by Glomeromycota and Mucoromycota, sister clades of the more well-known and diverse dikaryan fungi (all three are together called "symbiomycota"). AM fungi help plants to capture nutrients such as phosphorus, sulfur, nitrogen and micronutrients from the soil. It is believed that the development of the arbuscular mycorrhizal symbiosis played a crucial role in the initial colonisation of land by plants and in the evolution of the vascular plants. It has been said that it is quicker to list the plants that do not form endomycorrhizae than those that do. This symbiosis is a ...
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Polysporangiophyta
Polysporangiophytes, also called polysporangiates or formally Polysporangiophyta, are plants in which the spore-bearing generation ( sporophyte) has branching stems (axes) that bear sporangia. The name literally means 'many sporangia plant'. The clade includes all land plants (embryophytes) except for the bryophytes (liverworts, mosses and hornworts) whose sporophytes are normally unbranched, even if a few exceptional cases occur. While the definition is independent of the presence of vascular tissue, all living polysporangiophytes also have vascular tissue, i.e., are vascular plants or tracheophytes. Extinct polysporangiophytes are known that have no vascular tissue and so are not tracheophytes. Early polysporangiophytes History of discovery Paleobotanists distinguish between micro- and megafossils. Microfossils are primarily spores, either single or in groups. Megafossils are preserved parts of plants large enough to show structure, such as stem cross-sections or branching pa ...
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Marine Regression
A marine regression is a geological process occurring when areas of submerged seafloor are exposed above the sea level. The opposite event, marine transgression, occurs when flooding from the sea covers previously-exposed land. Evidence of marine regressions and transgressions occurs throughout the fossil record, and the fluctuations are thought to have caused or contributed to several mass extinctions, such as the Permian-Triassic extinction event (250 million years ago) and Cretaceous–Paleogene extinction event (66 Ma). During the Permian-Triassic extinction, the largest extinction event in the Earth's history, the global sea level fell 250 m (820 ft). A major regression could itself cause marine organisms in shallow seas to go extinct, but mass extinctions tend to involve both terrestrial and aquatic species, and it is harder to see how a marine regression could cause widespread extinctions of land animals. Regressions are, therefore, seen as correlates or symptoms of major ...
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Lau Event
The Lau event was the last of three relatively minor mass extinctions (the Ireviken, Mulde, and Lau events) during the Silurian period. It had a major effect on the conodont fauna, but barely scathed the graptolites, though they suffered an extinction very shortly thereafter termed the Kozlowskii event that some authors have suggested was coeval with the Lau event and only appears asynchronous due to taphonomic reasons. It coincided with a global low point in sea level caused by glacioeustasy and is closely followed by an excursion in geochemical isotopes in the ensuing late Ludfordian faunal stage and a change in depositional regime. Biological impact The Lau event started at the beginning of the late Ludfordian, a subdivision of the Ludlow stage, about . Its strata are best exposed in Gotland, Sweden, taking its name from the parish of Lau. Its base is set at the first extinction datum, in the Eke beds, and despite a scarcity of data, it is apparent that most major groups suf ...
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Ludfordian
In the geologic timescale, the Ludfordian is the upper of two chronostratigraphic stages within the Ludlow Series. Its age is the late Silurian Period, and within both the Palaeozoic Era and Phanerozoic Eon. The rocks assigned to the Ludfordian date to between 425.6 ± 0.9 Ma and 423.0 ± 2.3 Ma (million years ago). The Ludfordian Stage succeeds the Gorstian Stage and precedes the Pridoli Epoch. It is named for the village of Ludford in Shropshire, England. The GSSP for the Ludfordian is represented as a thin shale seam, coincident with the base of the Leintwardine Formation, overlying the Bringewood Formation in England. Paleoclimate The Lau event is a rapid pulse of cooling during the Ludfordian, about ; it is identified by a pulse of extinctions and oceanic changes. It is one of the series of fast sea-level and excursions in oxygen isotope ratios that signal fast switches between warm and cold climate states, characteristic of the Silurian climatic instability. The Lau ...
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Andean-Saharan Glaciation
The Andean-Saharan glaciation, also known as the Early Palaeozoic Icehouse, the Early Palaeozoic Ice Age, the Late Ordovician glaciation, the end-Ordovician glaciation, or the Hirnantian glaciation, occurred during the Paleozoic from approximately 460 Ma to around 420 Ma, during the Late Ordovician and the Silurian period. The major glaciation during this period, which was formerly thought only to consist of the Hirnantian glaciation itself, but has now been recognized as a longer, more gradual event that began as early as the Darriwilian, is widely considered to be the leading cause of the Ordovician-Silurian extinction event. Evidence of this glaciation can be seen in places such as Arabia, North Africa, South Africa, Brazil, Peru, Bolivia, Chile, Argentina, and Wyoming. More evidence derived from isotopic data is that during the Late Ordovician, tropical ocean temperatures were about 5 °C cooler than present day; this would have been a major factor that aided in the glacia ...
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Homerian
In the geologic timescale, the Homerian is an age of the Wenlock Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that is comprehended between 430.5 ± 0.7 Ma and 427.4 ± 0.5 Ma (million years ago), approximately. The Homerian Age succeeds the Sheinwoodian Age and precedes the Gorstian Age. The name comes from the small village of Homer, Shropshire near Much Wenlock. The defining lower boundary of Homerian rock layers (GSSP) is located within the Coalbrookdale Formation Coalbrookdale Formation, earlier known as Wenlock Shale or Wenlock Shale Formation and also referred to as Herefordshire Lagerstätte in palaeontology, is a fossil-rich deposit ('' Konservat-Lagerstätte'') in Powys and Herefordshire at the Engla ... of England. References Wenlock epoch Silurian geochronology {{geochronology-stub ...
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Bryophyte
The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in size and prefer moist habitats although they can survive in drier environments. The bryophytes consist of about 20,000 plant species. Bryophytes produce enclosed reproductive structures (gametangia and sporangia), but they do not produce flowers or seeds. They reproduce sexually by spores and asexually by fragmentation or the production of gemmae. Though bryophytes were considered a paraphyletic group in recent years, almost all of the most recent phylogenetic evidence supports the monophyly of this group, as originally classified by Wilhelm Schimper in 1879. The term ''bryophyte'' comes . Terminology The term "Bryophyta" was first suggested by Braun in 1864. G.M. Smith placed this group between Algae and Pteridophyta. Features The d ...
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Embryophyte
The Embryophyta (), or land plants, are the most familiar group of green plants that comprise vegetation on Earth. Embryophytes () have a common ancestor with green algae, having emerged within the Phragmoplastophyta clade of green algae as sister of the Zygnematophyceae. The Embryophyta consist of the bryophytes plus the polysporangiophytes. Living embryophytes therefore include hornworts, liverworts, mosses, lycophytes, ferns, gymnosperms and flowering plants. The land plants have diplobiontic life cycles and it is accepted now that they emerged from freshwater, multi-celled algae. The embryophytes are informally called land plants because they live primarily in terrestrial habitats (with exceptional members who evolved to live once again in aquatic habitats), while the related green algae are primarily aquatic. Embryophytes are complex multicellular eukaryotes with specialized reproductive organs. The name derives from their innovative characteristic of nurturing the ...
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