Schizeales
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Schizeales
Schizaeales is an order of ferns (class Polypodiopsida). Description While the three clades of Schizaeales are all well-distinguished from one another by numerous morphological characters, members of the order all have dimorphic fertile and sterile fronds and lack well-defined sori. Their sporangia have a horizontal annulus that lies below and completely encircles the top of the sporangium. Classification In the molecular phylogenetic classification of Smith et al. in 2006, the Schizaeales were placed in the leptosporangiate ferns, class Polypodiopsida. Three families, Anemiaceae, Lygodiaceae, and Schizaeaceae were recognized. The linear sequence of Christenhusz et al. (2011), intended for compatibility with the classification of Chase and Reveal (2009) which placed all land plants in Equisetopsida, reclassified Smith's Polypodiopsida as subclass Polypodiidae and placed the Schizaeales there, with the same three families. The classification of Christenhusz and Chase (2014) ...
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Schizaeaceae
Schizaeaceae is a family of ferns in the order Schizaeales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it includes only two genera. Alternatively, two families kept separate in PPG I, Lygodiaceae and Anemiaceae, may be included in Schizaeaceae (as the subfamilies Lygodioideae and Anemioideae) so that the family has four genera. In this approach, the Schizaeaceae of PPG I is treated as the subfamily Schizaeoideae. Species are mainly distributed in the tropics, but several are found in temperate regions in North America, South Africa, Australasia and Northeast Asia. The sporangia are borne on specialised pinnae, distinct from ordinary vegetative pinnae. The pinnae form small comb-like, pinnate structures on which the sporangia are formed. Genera In the narrow circumscription of the family, there are two genera: *'' Actinostachys'' Wall *'' Schizaea'' Sm. Two further genera are included in the broader circumscription: *''Anemia'' Sm. – otherwise placed i ...
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Annulus (botany)
{{Short description, A ring of specialized cells on the sporangium An annulus in botany is an arc or a ring of specialized cells on the sporangium. These cells are arranged in a single row, and are associated with the release or dispersal of spores. Ferns In leptosporangiate ferns, the annulus located on the outer rim of the sporangium and serves in spore dispersal. It consists typically of a ring or belt of dead water-filled cells with differentially thickened cell walls that stretches about two-thirds around each sporangium in leptosporangiate ferns. The thinner walls on the outside allow water to evaporate quickly under dry conditions. This dehiscence causes the cells to shrink and a contraction and straightening of the annulus ring, eventually rupturing the sporangial wall by ripping apart thin-walled lip cells on the opposite side of the sporangium. As more water evaporates, air bubbles form in the cells causing the contracted annulus to snap forward again, thus dislodging an ...
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Salviniales
The order Salviniales (formerly known as the Hydropteridales and including the former Marsileales) is an order of ferns in the class Polypodiopsida. Description Salviniales are all aquatic and differ from all other ferns in being heterosporous, meaning that they produce two different types of spores (megaspores and microspores) that develop into two different types of gametophytes (female and male gametophytes, respectively), and in that their gametophytes are endosporic, meaning that they never grow outside the spore wall and cannot become larger than the spores that produced them. The megasporangia each produce a single megaspore. In being heterosporus with endosporic gametophytes they are more similar to seed plants than to other ferns. The fertile and sterile leaves are dimorphic, taking on a different shape, and leaves bear anastomosing veins. Aerenchyma is frequently present in roots, shoots, and petioles (leaf stalks). The ferns of this order vary radically in form and ...
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Vine
A vine (Latin ''vīnea'' "grapevine", "vineyard", from ''vīnum'' "wine") is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas or runners. The word ''vine'' can also refer to such stems or runners themselves, for instance, when used in wicker work.Jackson; Benjamin; Daydon (1928). ''A Glossary of Botanic Terms with their Derivation and Accent'', 4th ed. London: Gerald Duckworth & Co. In parts of the world, including the British Isles, the term "vine" usually applies exclusively to grapevines (''Vitis''), while the term "climber" is used for all climbing plants. Growth forms Certain plants always grow as vines, while a few grow as vines only part of the time. For instance, poison ivy and bittersweet can grow as low shrubs when support is not available, but will become vines when support is available. A vine displays a growth form based on very long stems. This has two purposes. A vine may use rock exposures, other plants, or other ...
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Frond
A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the large leaves of cycads, as well as palms (Arecaceae) and various other flowering plants, such as mimosa or sumac. "Frond" is commonly used to identify a large, compound leaf, but if the term is used botanically to refer to the leaves of ferns and algae it may be applied to smaller and undivided leaves. Fronds have particular terms describing their components. Like all leaves, fronds usually have a stalk connecting them to the main stem. In botany, this leaf stalk is generally called a petiole, but in regard to fronds specifically it is called a stipe, and it supports a flattened blade (which may be called a lamina), and the continuation of the stipe into this portion is called the rachis. The blades may be simple (undivided), pinnatifid ( ...
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Polypodiidae (plant)
The Polypodiidae, commonly called leptosporangiate ferns, formerly Leptosporangiatae, are one of four subclasses of ferns, and the largest of these, being the largest group of living ferns, including some 11,000 species worldwide. The group has also been treated as the class Pteridopsida or Polypodiopsida, although other classifications assign them a different rank. Older names for the group include Filicidae and Filicales, although at least the "water ferns" (now the Salviniales) were then treated separately. The leptosporangiate ferns are one of the four major groups of ferns, with the other three being the eusporangiate ferns comprising the marattioid ferns (Marattiidae, Marattiaceae), the horsetails (Equisetiidae, Equisetaceae), and whisk ferns and moonworts. There are approximately 8465 species of living leptosporangiate ferns, compared with about 2070 for all other ferns, totalling 10535 species of ferns. Almost a third of leptosporangiate fern species are epiphytes. These ...
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Phytotaxa
''Phytotaxa'' is a peer-reviewed scientific journal for rapid publication on any aspect of systematic botany. It publishes on a wide range of subjects, but focuses on new species, monographs, floras, revisions, reviews, and typification issues. ''Phytotaxa'' covers all plant groups covered by the International Code of Nomenclature for algae, fungi, and plants, including diatoms, fungi, algae, lichens, mosses, hornworts, liverworts, and vascular plants), both living and fossil. The journal was established in 2009 by Maarten Christenhusz and the first issue appeared in October 2009. Authors have the option to publish open access. Abstracting and indexing The journal is abstracted and indexed in Science Citation Index Expanded, Current Contents/Agriculture, Biology & Environmental Sciences, and BIOSIS Previews. See also * Zootaxa ''Zootaxa'' is a peer-reviewed scientific mega journal for animal taxonomists. It is published by Magnolia Press (Auckland, New Zealand). The ...
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Polypodiopsida
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Leptosporangiate Ferns
The Polypodiidae, commonly called leptosporangiate ferns, formerly Leptosporangiatae, are one of four subclasses of ferns, and the largest of these, being the largest group of living ferns, including some 11,000 species worldwide. The group has also been treated as the class Pteridopsida or Polypodiopsida, although other classifications assign them a different rank. Older names for the group include Filicidae and Filicales, although at least the "water ferns" (now the Salviniales) were then treated separately. The leptosporangiate ferns are one of the four major groups of ferns, with the other three being the eusporangiate ferns comprising the marattioid ferns (Marattiidae, Marattiaceae), the horsetails (Equisetiidae, Equisetaceae), and whisk ferns and moonworts. There are approximately 8465 species of living leptosporangiate ferns, compared with about 2070 for all other ferns, totalling 10535 species of ferns. Almost a third of leptosporangiate fern species are epiphytes. These ...
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Sporangium
A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cycle. Sporangia can produce spores by mitosis, but in nearly all land plants and many fungi, sporangia are the site of meiosis and produce genetically distinct haploid spores. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains haploid nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and germinate into haploid hyphae. Although sexual reproduction in fungi varies between phyla, for some fungi the sporangium plays an indirect role in sexual reprod ...
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Anemiaceae
''Anemia'' is a genus of ferns. It is the only genus in the family Anemiaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the genus may be placed as the only genus in the subfamily Anemioideae of a more broadly defined family Schizaeaceae, the family placement used in ''Plants of the World Online'' . Its species are sometimes called flowering ferns, but this term is more commonly applied to ferns of the genus ''Osmunda''. Fronds are dimorphic; in fertile fronds, the two lowermost pinnae are highly modified to bear the sporangia. Ferns in this genus have chromosome numbers based on ''x''=38: ''n''=38, 76, 114. Taxonomy The genus ''Anemia'' was first described by the Swedish botanist Olof Swartz in 1806. The family Anemiaceae was created by Johann Link in 1841. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), ''Anemia'' includes ''Mohria'' and ''Colina'' and is the only genus in the family. Some sources do not separate the f ...
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Frond Dimorphism
Frond dimorphism refers to a difference in ferns between the fertile and sterile fronds. Since ferns, unlike flowering plants, bear spores on the leaf blade itself, this may affect the form of the frond itself. In some species of ferns, there is virtually no difference between the fertile and sterile fronds, such as in the genus ''Dryopteris'', other than the mere presence of the sori, or fruit-dots, on the back of the fronds. Some other species, such as ''Polystichum acrostichoides'' (Christmas fern), or some ferns of the genus ''Osmunda'', feature dimorphism on a portion of the frond only. Others, such as some species of ''Blechnum'' and ''Woodwardia'', have fertile fronds that are markedly taller than the sterile. Still others, such as ''Osmunda cinnamomea'' (Cinnamon fern), or plants of the family Onocleaceae, have fertile fronds that are completely different from the sterile. Only members of the ''Onocleaceae'' and ''Blechnaceae'' exhibit a propensity towards dimorphy, while ...
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