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Specialized Metabolism In Oryza
''Oryza'' is a genus in the plant family Poaceae, of which ''Oryza sativa'' (cultivated rice) is a member. ''Oryza'' is a genus of 24 species, most of which are annual and some perennial grasses, which are found in tropical and swampy parts of Africa, Asia and Australia. Given its wide geographic range, there exists a substantial diversity of secondary metabolites, specialized metabolites (also called secondary metabolites) in the genus ''Oryza''. Understanding this diversity can provide us solutions for mitigate crop losses due to disease and pest damage in rice, and boost agricultural production. Anthocyanins and flavonols * Cyanidin 3-O-glucoside (I): Produced by purple pigmented rice * Malvidin: Produced by purple pigmented rice * Tricin * Quercitin * Kaempferol * Apigenin Carotenoids- * β-carotene (provitamin A) * Lutein * zeaxanthin Hydroxycinnamic acids Alkaloids * Benzoxazinoids * Quinolone alkaloids Phytoalexins * Sakuranetin: Increases rapidly under biotic and abio ...
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Oryza
''Oryza'' is a genus of plants in the grass family. It includes the major food crop rice (species ''Oryza sativa'' and ''Oryza glaberrima''). Members of the genus grow as tall, wetland grasses, growing to tall; the genus includes both annual and perennial species. ''Oryza'' is situated in tribe Oryzeae, which is characterized morphologically by its single-flowered spikelets whose glumes are almost completely suppressed. In ''Oryza'', two sterile lemma simulate glumes. The tribe Oryzeae is in subfamily Ehrhartoideae, a group of Poaceae tribes with certain features of internal leaf anatomy in common. The most distinctive leaf characteristics of this subfamily are the arm cells and fusoid cells found in their leaves.Heywood, V.H. Flowering Plants of the World 1993 Oxford University Press One species, Asian rice ( ''O. sativa''), provides 20% of global grain and is a food crop of major global importance. The species are divided into two subgroups within the genus. Species Inside ...
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Naringenin
Naringenin is a flavorless, colorless flavanone, a type of flavonoid. It is the predominant flavanone in grapefruit, and is found in a variety of fruits and herbs. Structure Naringenin has the skeleton structure of a flavanone with three hydroxy groups at the 4', 5, and 7 carbons. It may be found both in the aglycol form, naringenin, or in its glycosidic form, naringin, which has the addition of the disaccharide neohesperidose attached via a glycosidic linkage at carbon 7. Like the majority of flavanones, naringenin has a single chiral center at carbon 2, although the optical purity is variable. Racemization of S(-)-naringenin has been shown to occur fairly quickly. Sources and bioavailability Naringenin and its glycoside has been found in a variety of herbs and fruits, including grapefruit, bergamot, sour orange, tart cherries, tomatoes, cocoa, Greek oregano, water mint, as well as in beans. Ratios of naringenin to naringin vary among sources, as do enantiomeric rati ...
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Phytochemistry
Phytochemistry is the study of phytochemicals, which are chemicals derived from plants. Phytochemists strive to describe the structures of the large number of secondary metabolites found in plants, the functions of these compounds in human and plant biology, and the biosynthesis of these compounds. Plants synthesize phytochemicals for many reasons, including to protect themselves against insect attacks and plant diseases. The compounds found in plants are of many kinds, but most can be grouped into four major biosynthetic classes: alkaloids, phenylpropanoids, polyketides, and terpenoids. Phytochemistry can be considered a subfield of botany or chemistry. Activities can be led in botanical gardens or in the wild with the aid of ethnobotany. Phytochemical studies directed toward human (i.e. drug discovery) use may fall under the discipline of pharmacognosy, whereas phytochemical studies focused on the ecological functions and evolution of phytochemicals likely fall under the disci ...
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Strigolactone
Strigolactones are a group of chemical compounds produced by a plant's roots. Due to their mechanism of action, these molecules have been classified as plant hormones or phytohormones. So far, strigolactones have been identified to be responsible for three different physiological processes: First, they promote the germination of parasitic organisms that grow in the host plant's roots, such as ''Striga'' ''lutea'' and other plants of the genus ''Striga''. Second, strigolactones are fundamental for the recognition of the plant by symbiotic fungi, especially arbuscular mycorrhizal fungi, because they establish a mutualistic association with these plants, and provide phosphate and other soil nutrients. Third, strigolactones have been identified as branching inhibition hormones in plants; when present, these compounds prevent excess bud growing in stem terminals, stopping the branching mechanism in plants. Strigolactones comprise a diverse group, but they all have core common chemical ...
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Brassinosteroid
Brassinosteroids (BRs or less commonly BS) are a class of polyhydroxysteroids that have been recognized as a sixth class of plant hormones and may have utility as an anticancer drug for endocrine-responsive cancers to induce apoptosis and inhibit growth. These brassinosteroids were first explored during the 70s, when Mitchell et al. reported promotion in stem elongation and cell division by the treatment of organic extracts of rapeseed ('' Brassica napus'') pollen. Brassinolide was the first isolated brassinosteroid in 1979, when pollen from '' Brassica napus'' was shown to promote stem elongation and cell divisions, and the biologically active molecule was isolated. The yield of brassinosteroids from 230 kg of ''Brassica napus'' pollen was only 10 mg. Since their discovery, over 70 BR compounds have been isolated from plants. Biosynthesis The BR is biosynthesised from campesterol. The biosynthetic pathway was elucidated by Japanese researchers and later shown to ...
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Abscisic Acid
Abscisic acid (ABA) is a plant hormone. ABA functions in many plant developmental processes, including seed and bud dormancy, the control of organ size and stomatal closure. It is especially important for plants in the response to environmental stresses, including drought, soil salinity, cold tolerance, freezing tolerance, heat stress and heavy metal ion tolerance. In plants Function ABA was originally believed to be involved in abscission, which is how it received its name. This is now known to be the case only in a small number of plants. ABA-mediated signaling also plays an important part in plant responses to environmental stress and plant pathogens. The plant genes for ABA biosynthesis and sequence of the pathway have been elucidated. ABA is also produced by some plant pathogenic fungi via a biosynthetic route different from ABA biosynthesis in plants. In preparation for winter, ABA is produced in terminal buds. This slows plant growth and directs leaf primordia to devel ...
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Gibberelin
Gibberellic acid (also called gibberellin A3, GA, and GA3) is a hormone found in plants and fungi. Its chemical formula is C19H22O6. When purified, it is a white to pale-yellow solid. Plants in their normal state produce large amounts of GA3. It is possible to produce the hormone industrially using microorganisms.Camara, M. C. et al (2015) General Aspects and Applications of Gibberelins and Gibberellic Acid in Plants. In: Hardy, J.. (Org.). Gibberellins and Gibberellic Acid: Biosynthesis, Regulation and Physiological Effects. 1ed.Hauppauge: Nova Science Publishers, 2015, v., p. 1-21. Gibberellic acid is a simple gibberellin, a pentacyclic diterpene acid promoting growth and elongation of cells. It affects decomposition of plants and helps plants grow if used in small amounts, but eventually plants develop tolerance to it. GA stimulates the cells of germinating seeds to produce mRNA molecules that code for hydrolytic enzymes. Gibberellic acid is a very potent hormone whose natural ...
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Cytokinin
Cytokinins (CK) are a class of plant hormones that promote cell division, or cytokinesis, in plant roots and shoots. They are involved primarily in cell growth and differentiation, but also affect apical dominance, axillary bud growth, and leaf senescence. There are two types of cytokinins: adenine-type cytokinins represented by kinetin, zeatin, and 6-benzylaminopurine, and phenylurea-type cytokinins like diphenylurea and thidiazuron (TDZ). Most adenine-type cytokinins are synthesized in roots. Cambium and other actively dividing tissues also synthesize cytokinins. No phenylurea cytokinins have been found in plants. Cytokinins participate in local and long-distance signalling, with the same transport mechanism as purines and nucleosides. Typically, cytokinins are transported in the xylem. Cytokinins act in concert with auxin, another plant growth hormone. The two are complementary, having generally opposite effects. History The idea of specific substances required for ...
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Auxin
Auxins (plural of auxin ) are a class of plant hormones (or plant-growth regulators) with some morphogen-like characteristics. Auxins play a cardinal role in coordination of many growth and behavioral processes in plant life cycles and are essential for plant body development. The Dutch biologist Frits Warmolt Went first described auxins and their role in plant growth in the 1920s. Kenneth V. Thimann became the first to isolate one of these phytohormones and to determine its chemical structure as indole-3-acetic acid (IAA). Went and Thimann co-authored a book on plant hormones, ''Phytohormones'', in 1937. Overview Auxins were the first of the major plant hormones to be discovered. They derive their name from the Greek word αυξειν (''auxein'' – "to grow/increase"). Auxin is present in all parts of a plant, although in very different concentrations. The concentration in each position is crucial developmental information, so it is subject to tight regulation through both meta ...
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Jasmonate
Jasmonate (JA) and its derivatives are lipid-based plant hormones that regulate a wide range of processes in plants, ranging from growth and photosynthesis to reproductive development. In particular, JAs are critical for plant defense against herbivory and plant responses to poor environmental conditions and other kinds of abiotic and biotic challenges. Some JAs can also be released as volatile organic compounds (VOCs) to permit communication between plants in anticipation of mutual dangers. History The isolation of methyl jasmonate (MeJa) from jasmine oil derived from ''Jasminum grandiflorum'' led to the discovery of the molecular structure of jasmonates and their name in 1962 while jasmonic acid itself was isolated from ''Lasiodiplodia theobromae'' by Alderidge et al in 1971. Biosynthesis Biosynthesis is reviewed by Acosta and Farmer 2010, Wasternack and Hause 2013, and Wasternack and Song 2017. Jasmonates (JA) are oxylipins, i.e. derivatives of oxygenated fatty acid. The ...
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Salicylate
Salicylic acid is an organic compound with the formula HOC6H4CO2H. A colorless, bitter-tasting solid, it is a precursor to and a metabolite of aspirin (acetylsalicylic acid). It is a plant hormone, and has been listed by the EPA Toxic Substances Control Act (TSCA) Chemical Substance Inventory as an experimental teratogen. The name is from Latin ''salix'' for willow tree. It is an ingredient in some anti-acne products. Salts and esters of salicylic acid are known as salicylates. Uses Medicine Salicylic acid as a medication is commonly used to remove the outer layer of the skin. As such, it is used to treat warts, psoriasis, acne vulgaris, ringworm, dandruff, and ichthyosis. Similar to other hydroxy acids, salicylic acid is an ingredient in many skincare products for the treatment of seborrhoeic dermatitis, acne, psoriasis, calluses, corns, keratosis pilaris, acanthosis nigricans, ichthyosis, and warts. Uses in manufacturing Salicylic acid is used as a food preservativ ...
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Ethylene
Ethylene (IUPAC name: ethene) is a hydrocarbon which has the formula or . It is a colourless, flammable gas with a faint "sweet and musky" odour when pure. It is the simplest alkene (a hydrocarbon with carbon-carbon double bonds). Ethylene is widely used in the chemical industry, and its worldwide production (over 150 million tonnes in 2016) exceeds that of any other organic compound. Much of this production goes toward polyethylene, a widely used plastic containing polymer chains of ethylene units in various chain lengths. Ethylene is also an important natural plant hormone and is used in agriculture to force the ripening of fruits. The hydrate of ethylene is ethanol. Structure and properties This hydrocarbon has four hydrogen atoms bound to a pair of carbon atoms that are connected by a double bond. All six atoms that comprise ethylene are coplanar. The H-C-H angle is 117.4°, close to the 120° for ideal sp² hybridized carbon. The molecule is also relatively weak: rota ...
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