Pteraspidomorphi
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Pteraspidomorphi
Pteraspidomorphi is an extinct class of early jawless fish. They have long been regarded as closely related or even ancestral to jawed vertebrates, but the few characteristics they share with the latter are now considered as basal traits for all vertebrates. Characteristics Pteraspidomorphs are characterized by their massive dermal head armour having large, median, ventral and dorsal plates or shields. Janvier, Philippe (1997Pteraspidomorphi''The Tree of Life Web Project''. The fossils show extensive shielding of the head. Many had hypocercal tails in order to generate lift to increase ease of movement through the water for their armoured bodies, which were covered in dermal bone. They also had sucking mouth parts and some species may have lived in fresh water. Most pteraspidomorphs were marine, but lived very near to the shore, in lagoons and deltas. Some groups are thought to have been fresh water-dwelling. They were certainly bottom-dwellers, as shown by traces of abrasi ...
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Pteraspidomorphi
Pteraspidomorphi is an extinct class of early jawless fish. They have long been regarded as closely related or even ancestral to jawed vertebrates, but the few characteristics they share with the latter are now considered as basal traits for all vertebrates. Characteristics Pteraspidomorphs are characterized by their massive dermal head armour having large, median, ventral and dorsal plates or shields. Janvier, Philippe (1997Pteraspidomorphi''The Tree of Life Web Project''. The fossils show extensive shielding of the head. Many had hypocercal tails in order to generate lift to increase ease of movement through the water for their armoured bodies, which were covered in dermal bone. They also had sucking mouth parts and some species may have lived in fresh water. Most pteraspidomorphs were marine, but lived very near to the shore, in lagoons and deltas. Some groups are thought to have been fresh water-dwelling. They were certainly bottom-dwellers, as shown by traces of abrasi ...
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Eriptychiida
Eriptychiida is an extinct marine taxon of vertebrate in the group Pteraspidomorphi. The order contains the genus, '' Eriptychius'', and fossilized specimens from this genus have been found in the Gull River Formation of Ontario, the Harding Formation of Colorado, and the Bighorn Dolomite of Wyoming. The group contains two documented species: ''Eriptychius americanus'' and ''Eriptychius orvigi''. Characteristics The structure of the dentine of eriptychiids is in many respects closer to that of heterostracans that to that of astraspids. This is the only argument to place them as the closest relatives to heterostracans, among the Ordovician vertebrates. However, eriptychiids differ from all other pteraspidomorphs in having a massively calcified endoskeleton, pervaded by canals for blood vessels. Taxonomy * Order † Eriptychiiformes Ørvig 1958 ** Genus ?†'' Eleochera'' Sansom & Smith 2005 ** Family † Eriptychiidae Tarlo 1962 *** Genus †'' Eriptychius'' Walcott 1892 * ...
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Arandaspidiformes
Arandaspida is a taxon of very early, jawless prehistoric fish which lived during the Ordovician period. Arandaspids represent the oldest known craniates, a proposed group of chordates that contain all chordates with a cartilage-derived skull (i.e., lampreys, armored agnathans, and gnathostomes), and hagfish. The group represents a subclass within the class Pteraspidomorphi, and contains only one order, the Arandaspidiformes. The oldest known genus of this group is ''Sacabambaspis'' found in South America. Characteristics The head armor of arandaspids is elongated, fusiform, with a rather flat dorsal shield, and a bulging ventral shield. In the anterior part of the dorsal shield are two closely set holes, which have been thought to be a paired pineal opening, but which are more likely the external openings of the endolymphatic ducts. The eyes, surrounded by a sclerotic ring, are housed in a notch at the anterior end of the dorsal shield. The nostrils are not clearly located, b ...
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Jawless Fish
Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present (cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animals, cyclostomes are sister to all vertebrates with jaws, known as gnathostomes. Recent molecular data, both from rRNA and from mtDNA as well as embryological data, strongly supports the hypothesis that living agnathans, the cyclostomes, are monophyletic. The oldest fossil agnathans appeared in the Cambrian, and two groups still survive today: the lampreys and the hagfish, comprising about 120 species in total. Hagfish are considered members of the subphylum Vertebrata, because they secondarily lost vertebrae; before this event was inferred from molecular and developmental data, the group Craniata was created by Linnaeus (and is still sometimes used as a strictly morphological descriptor) to reference hagfish plus vertebrates. While a few scientist ...
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Arandaspida
Arandaspida is a taxon of very early, jawless prehistoric fish which lived during the Ordovician period. Arandaspids represent the oldest known craniates, a proposed group of chordates that contain all chordates with a cartilage-derived skull (i.e., lampreys, armored agnathans, and gnathostomes), and hagfish. The group represents a subclass within the class Pteraspidomorphi, and contains only one order, the Arandaspidiformes. The oldest known genus of this group is ''Sacabambaspis'' found in South America. Characteristics The head armor of arandaspids is elongated, fusiform, with a rather flat dorsal shield, and a bulging ventral shield. In the anterior part of the dorsal shield are two closely set holes, which have been thought to be a paired pineal opening, but which are more likely the external openings of the endolymphatic ducts. The eyes, surrounded by a sclerotic ring, are housed in a notch at the anterior end of the dorsal shield. The nostrils are not clearly located, b ...
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Astraspida
Astraspida, or astraspids, are a small group of extinct armored jawless vertebrates, which lived in the Middle Ordovician (about 450 million years ago) in North America. They are placed among the Pteraspidomorphi because of the large dorsal and ventral shield of their head armor. They are represented by a single genus, Astraspis, including possibly two species, ''A. desiderata'' and ''A. splendens'' but their remains are fairly abundant in Ordovician sandstones of the USA (Colorado, Arizona, Oklahoma, Wyoming) and Canada (Quebec). The head armor of Astraspis is rather massive, with a series of ten gill openings lining the margin of the dorsal shield, and laterally placed eyes. The dorsal shield is ribbed by strong longitudinal crests, and the tail is covered with large, diamond-shaped scales. They are often grouped together with the Arandaspidida. Characteristics Astraspids are characterized by a dermal ornamentation of large, mushroom shaped tubercles of fine-tubuled dentine ("ast ...
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Heterostraci
Heterostraci (Ancient Greek, ἕτερος+ὄστρακον "those itha different shell" i is pl. of -us is an extinct subclass of pteraspidomorph jawless vertebrate that lived primarily in marine and estuary environments. Heterostraci existed from the mid-Ordovician to the conclusion of the Devonian. Description and anatomy The Heterostracans differed from other Paleozoic agnathan taxa both in the arrangement and histology of their scales. Most heterostracans had two plates which form a large dorsal shield and a large ventral shield, and had series of scales arranged in various patterns on the sides of their bodies, the exact pattern differing from one group to another. In a few primitive forms, such as '' Lepidaspis'', the dorsal and ventral shields are composed of a mosaic of tiny scales. In most other known forms, though, these tiny scales have fused together to form the shield-plates. The scales of heterostracans are histologically distinct from other vertebrate ...
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Vertebrates
Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with currently about 69,963 species described. Vertebrates comprise such groups as the following: * jawless fish, which include hagfish and lampreys * jawed vertebrates, which include: ** cartilaginous fish (sharks, rays, and ratfish) ** bony vertebrates, which include: *** ray-fins (the majority of living bony fish) *** lobe-fins, which include: **** coelacanths and lungfish **** tetrapods (limbed vertebrates) Extant vertebrates range in size from the frog species ''Paedophryne amauensis'', at as little as , to the blue whale, at up to . Vertebrates make up less than five percent of all described animal species; the rest are invertebrates, which lack vertebral columns. The vertebrates traditionally include the hagfish, which do no ...
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Ostracoderm
Ostracoderms () are the armored jawless fish of the Paleozoic Era. The term does not often appear in classifications today because it is paraphyletic (excluding jawed fishes) (may also be polyphyletic if anaspids are closer to cyclostomes) and thus does not correspond to one evolutionary lineage. However, the term is still used as an informal way of loosely grouping together the armored jawless fishes. An innovation of ostracoderms was the use of gills not for feeding, but exclusively for respiration. Earlier chordates with gill precursors used them for both respiration and feeding. Ostracoderms had separate pharyngeal gill pouches along the side of the head, which were permanently open with no protective operculum. Unlike invertebrates that use ciliated motion to move food, ostracoderms used their muscular pharynx to create a suction that pulled small and slow moving prey into their mouths. Swiss anatomist Louis Agassiz received some fossils of bony armored fish from Scotland ...
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Ostracoderm
Ostracoderms () are the armored jawless fish of the Paleozoic Era. The term does not often appear in classifications today because it is paraphyletic (excluding jawed fishes) (may also be polyphyletic if anaspids are closer to cyclostomes) and thus does not correspond to one evolutionary lineage. However, the term is still used as an informal way of loosely grouping together the armored jawless fishes. An innovation of ostracoderms was the use of gills not for feeding, but exclusively for respiration. Earlier chordates with gill precursors used them for both respiration and feeding. Ostracoderms had separate pharyngeal gill pouches along the side of the head, which were permanently open with no protective operculum. Unlike invertebrates that use ciliated motion to move food, ostracoderms used their muscular pharynx to create a suction that pulled small and slow moving prey into their mouths. Swiss anatomist Louis Agassiz received some fossils of bony armored fish from Scotland ...
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Cyathaspidiformes
Cyathaspidiformes is an extinct order of heterostracan vertebrates known from extensive fossil remains primarily from Silurian to Early Devonian strata of Europe, and North America, and from Early Devonian marine strata of Siberia. Anatomy Like their descendants, the pteraspidids, all cyathaspidiform heterostracans had the cephalothorax enclosed in armor, formed from several plates, including dorsal, ventral, a dorsal spine derived from a scale, and a large, scale-covered tail. Thus, the living animals would have resembled tadpoles encased in massive armor. The majority of taxa have the rostral and pineal plates fused or merged with the dorsal plate, and in the amphiaspidids, all the plates of the cephalothorax were fused together into a single "muff-like" unit. Unlike the pteraspidids, all cyathaspidiforms are thought to be almost uniformly benthic in lifestyle, though only the amphiaspids and the ctenaspids are thought to be burrowers. Taxonomy The taxonomy of Cyathaspidifo ...
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Pteraspis Rostrata
''Pteraspis'' (from el, πτερόν 'wing' or 'fin' and el, ἀσπίς 'shield') is an extinct genus of pteraspidid heterostracan agnathan vertebrate that lived from the Lochkovian to Eifelian epochs of the Devonian period in what is now Brazil (Eifelian Maecuru Formation), Britain (Lochkovian Ditton Group), Ukraine (Lochkovian Ivane Suite, Pragian Babin Sandstone) and Belgium. Description Like other heterostracan fishes, ''Pteraspis'' had a protective armored plating covering the front of its body. Though lacking fins other than its lobed tail, it is thought to have been a good swimmer thanks to stiff, wing-like protrusions derived from the armoured plates over its gills. This, along with the horn-like rostrum, made ''Pteraspis'' very streamlined in shape; a perfect quality for a good swimmer. ''Pteraspis'' also had some stiff spikes on its back, possibly an additional form of protection against predators. It is thought to have fed from shoals of plankton Pla ...
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