Protungulatum
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Protungulatum
''Protungulatum'' ('first ungulate') is a extinct genus of pan-euungulate mammals within extinct family Protungulatidae, and also one of the earliest known placental mammals in the fossil record, that lived in North America from the Late Cretaceous to early Paleocene.Maeva Orliac (2016."The inner ear of ''Protungulatum'' (Pan-Euungulata, Mammalia)"Journal of Mammalian Evolution 23(4) Fossils of this genus were first found in the Bug Creek Anthills in northeastern Montana. The Bug Creek Anthills were initially believed to be Late Cretaceous (latest Maastrichtian) because of the presence of the remains of non-avian dinosaurs and common Cretaceous mammals, but these were later shown to have been reworked from Late Cretaceous strata, and consequently the Bug Creek Anthills are currently believed to be Early Paleocene (Puercan) in age. Remains from the Ravenscrag Formation of Saskatchewan, Canada have been assigned to ''Protungulatum donnae''. These remains may also be Cretaceous ...
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Ungulate
Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. The term means, roughly, "being hoofed" or "hoofed animal". As a descriptive term, "ungulate" normally excludes cetaceans as they do not possess most of the typical morphological characteristics of other ungulates, but recent discoveries indicate that they were also descended from early artiodactyls. Ungulates are typically herbivorous and many employ specialized gut bacteria to allow them to digest cellulose. Some modern species, such as pigs, are omnivorous, ...
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Euungulata
Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. The term means, roughly, "being hoofed" or "hoofed animal". As a descriptive term, "ungulate" normally excludes cetaceans as they do not possess most of the typical morphological characteristics of other ungulates, but recent discoveries indicate that they were also descended from early artiodactyls. Ungulates are typically herbivorous and many employ specialized gut bacteria to allow them to digest cellulose. Some modern species, such as pigs, are omnivorous, w ...
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Condylarthra
Condylarthra is an informal group – previously considered an order – of extinct placental mammals, known primarily from the Paleocene and Eocene epochs. They are considered early, primitive ungulates. It is now largely considered to be a wastebasket taxon, having served as a dumping ground for classifying ungulates which had not been clearly established as part of either Perissodactyla or Cetartiodactyla, being composed thus of several unrelated lineages. Taxonomic history Condylarthra always was a problematic group. When Condylarthra was first described by , Phenacodontidae was the type and only family therein. , however, raised Condylarthra to an order and included a wide range of diverse placentals with generalized dentitions and postcranial skeletons. More recent researchers (i.e. post-WW2) have been more restrictive; either including only a limited number of taxa, or proposing that the term should be abandoned altogether. Due to their primitive characteristics condylar ...
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Placentalia
Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguished from monotremes and marsupials in that the fetus (biology), fetus is carried in the uterus of its mother to a relatively late stage of development. The name is something of a misnomer considering that marsupials also nourish their fetuses via a placenta, though for a relatively briefer period, giving birth to less developed young which are then nurtured for a period inside the mother's pouch (marsupial), pouch. Anatomical features Placental mammals are anatomically distinguished from other mammals by: * a sufficiently wide opening at the bottom of the pelvis to allow the birth of a large baby relative to the size of the mother. * the absence of epipubic bones extending forward from the pelvis, which are found in all other mammals. (Their ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saur ...
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Arctocyonidae
Arctocyonidae (from Greek ''arktos'' ''kyôn'', "bear/dog-like") has been defined as an extinct family of unspecialized, primitive mammals with more than 20 genera. Animals assigned to this family were most abundant during the Paleocene, but extant from the late Cretaceous to the early Eocene (). Like most early mammals, their actual relationships are very difficult to resolve. No Paleocene fossil has been unambiguously assigned to any living order of placental mammals, and many genera resemble each other: generalized robust, not very agile animals with long tails and all-purpose chewing teeth, living in warm closed-canopy forests with many niches left vacant by the K-T extinction. Arctocyonids were early defined as a family of creodonts (early predators), then reassigned to the condylarths (primitive plant-eaters, now understood as a wastebasket taxon). More recently, these animals have been thought to be the ancestors of the orders Mesonychia and Cetartiodactyla, although some ...
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Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary extinction event) occurred at the end of this age. In this mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous. Stratigraphic definitions Definition The Maastrichtian was introduced into scientific literature by Belgian geologist André Hubert Dumont in 1849, after studying rock strata of the Chalk Group c ...
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Ravenscrag Formation
The Ravenscrag Formation is a stratigraphic unit of early Paleocene age in the Western Canada Sedimentary Basin. It was named for the settlement of Ravenscrag, Saskatchewan, and was first described from outcrops at Ravenscrag Butte near the Frenchman River by N.B. Davis in 1918.Davis, N.B., 1918. Report on the clay resources of southern Saskatchewan; Canada, Department of Mines, Mines Branch, Report 468, 93 p. The Ravenscrag Formation includes a wide range of vertebrate and plant fossils, as well as economically significant coal seams and clay deposits. Lithology The Ravenscrag Formation is an eastward-thickening wedge of sediments. It is composed primarily of buff, grey and white silty claystone, with mudstone, siltstone, sandstone, and several significant coal seams. Glass, D.J., editor, 1997. Lexicon of Canadian Stratigraphy, vol. 4, Western Canada, p. 974. Canadian Society of Petroleum Geologists, Calgary, Alberta, 1423 p. on CD-ROM, . These sediments were deposited in flood ...
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Scrotifera
Scrotifera ("beasts with scrotum") is a clade of Placentalia, placental mammals that groups together grandorder Ferungulata, clade Bat, Apo-Chiroptera (aka Order (biology), order Bat, Chiroptera), other extinct members and their common ancestors. The clade Scrotifera is a sister group to the order Eulipotyphla (true insectivores) based on evidence from molecular phylogenetics, and together they make superorder Laurasiatheria. The last common ancestor of Scrotifera is supposed to have diversified ca. 73.1 to 85.5 million years ago. Etymology Peter Waddell, then of the Institute of Statistical Mathematics, explains the etymology of the clade's name as follows: The name comes from the word ''scrotum'', a pouch in which the Testicle, testes permanently reside in the adult male. All members of the group have a postpenile scrotum, often prominently displayed, except for some aquatic forms and pangolin (which has the testes just below the skin). It appears to be an ancestral character ...
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Science (journal)
''Science'', also widely referred to as ''Science Magazine'', is the peer-reviewed academic journal of the American Association for the Advancement of Science (AAAS) and one of the world's top academic journals. It was first published in 1880, is currently circulated weekly and has a subscriber base of around 130,000. Because institutional subscriptions and online access serve a larger audience, its estimated readership is over 400,000 people. ''Science'' is based in Washington, D.C., United States, with a second office in Cambridge, UK. Contents The major focus of the journal is publishing important original scientific research and research reviews, but ''Science'' also publishes science-related news, opinions on science policy and other matters of interest to scientists and others who are concerned with the wide implications of science and technology. Unlike most scientific journals, which focus on a specific field, ''Science'' and its rival ''Nature (journal), Nature'' c ...
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Eutheria
Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the clade consisting of all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from noneutherians by various phenotypic traits of the feet, ankles, jaws and teeth. All extant eutherians lack epipubic bones, which are present in all other living mammals (marsupials and monotremes). This allows for expansion of the abdomen during pregnancy. The oldest-known eutherian species is '' Juramaia sinensis'', dated at from the early Late Jurassic ( Oxfordian) of China. Eutheria was named in 1872 by Theodore Gill; in 1880 Thomas Henry Huxley defined it to encompass a more broadly defined group than Placentalia. Characteristics Distinguishing features are: *an enlarged malleolus ("little hammer") at the bottom of the tibia, the larger of the two shin bones *the joint between the first metatarsal bone and the entocuneiform bone (the innermost of the three cuneiform ...
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Polyphyletic
A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of convergent evolution. The arrangement of the members of a polyphyletic group is called a polyphyly .. ource for pronunciation./ref> It is contrasted with monophyly and paraphyly. For example, the biological characteristic of warm-bloodedness evolved separately in the ancestors of mammals and the ancestors of birds; "warm-blooded animals" is therefore a polyphyletic grouping. Other examples of polyphyletic groups are algae, C4 photosynthetic plants, and edentates. Many taxonomists aim to avoid homoplasies in grouping taxa together, with a goal to identify and eliminate groups that are found to be polyphyletic. This is often the stimulus for major revisions of the classification schemes. Researchers concerned more with ecology than with systema ...
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